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What does it take to evolve behaviorally complex organisms?

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Figure

Figure 2. Modular and nonmodular architectures for the What and Where tasks.
Figure 3. Average fitness for the What and Where tasks across 50.000 generations for modular and  nonmodular architectures when network architecture is fixed and what evolves is only the set of connection weights for the architecture
Figure 4. Genetic interference. Conflicting mutations (one favorable and the other one unfavorable) can fall on separate portions of the genotype encoding distinct neural modules, thereby hindering the efficiency of the selection process
Figure 5. Average fitness (error) across 50.000 generations  when the genotype encodes both the Where and What tasks (single population) and when it encodes either one or the other of the two tasks (two populations)
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