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(1)

ISSN: 1992-8645 www.jatit.org E-ISSN: 1817-3195

EXPONENTIAL STABILITY OF STRONG SOLUTION FOR A

STOCHASTIC DISTURBANCE PREDATOR-PREY SYSTEM

OF THREE SPECIES WITH AGE-STRUCTURE

1CHENLIYU,2LIUSI

1Ningxia University,Yinchuan,750021People’s Republic of China 2

Zhejiang University ,Hangzhou,310058,People’s Republic of China Email: [email protected] , [email protected]

ABSTRACT

This thesis first introduces stochastic disturbance factor into a model and has a discussion of it on the basis of a predator-prey system of three species with age-structure proposed by Zhixue Luo. Sufficient conditions of a stochastic disturbance predator-prey system of three species with age-structure have been given and exponential stability of strong solution has been proved according to Gronwall’s lemma and

Ito

ˆ

formula. The conclusion is a further development of pertinent literature.

Keywords:Exponential Stability, Stochastic Disturbance,

Ito

ˆ

Formula, Gronwall’s Lemma, Three Species Subject Classification:(AMS) 35B10, 45D05, 60H15, 34K40

1. INTRODUCTION

The study of age-structured single species was initiated by Rorres and Fair[1]. Since then, the control problem has received many attentions from several authors[2-9]. In [2], Chan and Guo studied optimal birth control policies for the model of Mckendrick type. Maximum principles for problems with free ends, the time optimalcontrol problem, problems with target sects and infinite horizon problems had beenderived respectively. Motivated by the idea of Chan and Gou[2], the aim of this paperis to establish necessary optimality

conditions for the above mentioned optimalcontrol problems by using a powerful functional approach first suggested for generalexternal problems.In [10], Webb studied the stability of nontrivial equilibrium solution of the model.In [11], Zhixue Luo, Ze-Rong He and Wan-Tong Li considered optimal birth control for predator-prey system of three species with age-structure. Considering the effect of age factor for control problems of the interacting species. They introduce the following food chain system composed of three age-dependent species.

where

Q

=

(0,

a

+

) (0,

×

+∞

),[ ,

a a

1 2

]

is the

fertility interval , and the other parameters mean as follow: (for the sake of convenience, throughout this paper we suppose that i=1,2,3).

( , )

i

p a t

: the density of ith population of age a at

time t,

p

i

: , ;

=

p a t

i

( )

;

( , )

i

a t

µ

: the average mortality of ith population;

( )

i

t

β

: the average fertility of ith population;

( , )

k

a t

(2)

( , )

i

m a t

: the ratio of females in ith population;

0

( )

i

p

a

: the initial of age distribution of ith population;

a

+ : the life expectancy,

0

<

a

+

< +∞

. Here without loss of generality.

we assume that the three population have the same life expectancy.

There has been much recent interest in a three-species Predator-Prey System. For example, Tan DJ and Zhanp predator-prey system with impulsive perturbations on predators, Liu ZJ[13] researched existence of periodic solutions for a delayed ratiod-ependent three-species predator-prey diffusion system on time scales, In [14], Xu R and Chen LS discussed persistence and global stability for a three-species ratio-dependent predator-prey system with time delays in two-patch environments, In [15],

Zhang YP and Sun JT studied persistence in a three

species Lotka-Volterra non-periodic predator-prey system, Dong LZ, Yuan CD and Chen LS[16] discussed persistence of a three-species predator-prey-chain autonomous system with diffusion.

This thesis first introduces stochastic disturbance to a predator-prey system of three species with age-structure.

Suppose that in Eq(1), add

f p t dw

1

(

1

, )

to

1

( , )

a t p

1 1

( , )

a t P t p

2

( )

1

,

µ

λ

add

f

2

(

p t dw

2

, )

to−

µ

2( , )a t p2+

λ

2( , ) ( )a t P t p1 2

λ

3( , ) ( )a t P t p3 2,ad d

f p t dw

3

(

3

, )

to

µ

3

( , )

a t p

3

+

λ

4

( , )

a t P t p

2

( )

3 , where

ω

( )

t

is white noise. Then this environmentally perturbed system may be described by this system of three species with age-structure:

where

1( 1, )

f p t dw ,

f

2

(

p t dw

2

, )

,

f p t dw

3

(

3

, )

denote stochastically perturbed,

ω

( )

t

is white noise.

A new stochastic differential equation model (2) for a predator-prey system of three species with age-structure is derived. It is an extension of Eq. (1).

In this paper, we shall discussion the exponential stability of strong solution for stochastic predator-prey system of three species with age structure.

In Section 2, we begin with some preliminary results which are essential for our analysis. In Section 3, we shall establish some criteria for the exponential stability of strong solution for stochastic predator-prey system of three species with age structure.

2. PRELIMINARIES

Let

V

=

H

1

([0,

a

+

])

=

{ |

ϕ ϕ

L

2

([0,

a

+

]),

2

([0,

]),

L

a

a

ϕ

+

Where a

ϕ

∂ are generalized

partial derivatives

}

,

V

is the Sobolev

space.

H

=

L

2

([0,

a

+

])

such that

.

V

H

H

V

V

′is the dual space of

V

.We denote by

,

and

the norms in

V

,

H,and

V

′ respectively; by

⋅ ⋅

,

the duality product

between

V

,

V

′ and

⋅ ⋅

,

the scalar product in

H

, and m a constant such

that

| |

x

m x

‖ ‖

,

∀ ∈

x V

. For an operator

(

,

)

(3)

ISSN: 1992-8645 www.jatit.org E-ISSN: 1817-3195

operators from

M

into

H

,

‖ ‖

B

2 denotes the Hilbert-Schmidt norm, i.e. 2

2 ( )

T

B =tr BWB

‖ ‖ . Let

t

ω

be a Wiener process defined on complete probability space

( , , )

F P

and taking its values in the separable Hilbert space K,with increment convariance operator .Throughout this paper, we always assume that:

(H1)

µ

i

L Q

( ),

0 0

0

µ

i

µ

i

( , )

a t

µ

i

,

0 0

,

i i

µ µ

are constants,

0

( ,

)

, ( , )

a

i

a t

a da

a t

Q

µ

+

+

= +∞

;

(H2)

0

B

k

λ

k

( , )

a t

A

k

, ( , )

a t

Q A B

,

k

,

k

are constants, (k=1,2,3,4);

(H3)

0

m a t

i

( , )

M

i

, ( , )

a t

Q

,

M

i are

constants, and

m a t

i

( , )

0

, when

a

<

a

1

or

a

>

a

2;

(H4)

β

i Ui: {hi L (0, ) : 0

β

i0 h ti( )

β

i0, ∞

∈ = ∈ ∞ ≤ ≤ ≤

0}

t

∀ >

,

U

=

U

1

×

U

2

×

U

3.

(H5)

p

i0

L

(0,

a

),

p

i0

( )

a

0,

a

(0,

a

)

+ +

≥ ∀ ∈

;

(H6)

r

i

P t

i

( )

R P

i

,

i

:

=

P t

i

( ).

For any given

0

T

>

and 2 3

1 2 3

(

,

,

)

(0, ;

(0,

;

))

p

=

p p p

C

T L

a R

+

3

( T; )

L Q R

, where

Q

T

=

(0,

a

+

) (0, )

×

T

; (H7)Lipschitz:

2 3 3

(0, ;

(0,

;

))

(

;

),

i T

p

C

T L

a R

+

L Q R

∀ ∈

0,

i

K

∃ > such that

‖ ‖

f

i 22

K

i

|

p

i

| ,

2

i

=

1, 2, 3

,

For any given

T

>

0

and

2 3

1 2 3

(

,

,

)

(0, ;

(0,

;

))

p

=

p p p

C

T L

a R

+

3

( T; )

L Q R

Q

T

=

(0,

a

+

) (0, )

×

T

,

where

Q

T

=

(0,

a

+

) (0, )

×

T

.

3. EXPONENTIAL STABILITY OF SOLUTIONS

In this section, we shall establish some criteria for the exponential stability of stochastic competitive population system Eq(2). we assume there exists aprocess

p

=

(

p p p

1

,

2

,

3

)

2 3

(0, ;

(0,

;

))

C

T L

a R

+

3

(

T

;

)

L Q R

,

which is the strong solution of Eq(2).

Theorem3.1 Assume the preceding hypotheses

and 0 2 2 2

0

2Br+2

µ

−(

β

) M a+−2AR− >K 0 hold,

if

(

p p p

1

,

2

,

3

)

is a global strong solution to Eq(2).

There exist contants 0 2 2 2

0

2Br 2 ( ) M a

τ

= +

µ

β

+

2

AR

K

0,

C

0

− >

>

such that

2 2 2

1 2 3

(|

|

|

|

|

| )

t

,

0

E

p

+

p

+

p

Ce

−τ

∀ ≥

t

.

Proof. We can choose

δ

>

0

small enough such that

ξ δ

− >

0

. Then, by

Ito

ˆ

formula , we have

( ) 2 2 2

1 2 3

2 2 2

10 20 30

(|

|

|

|

|

| )

(|

|

|

|

|

| )

t

e

p

p

p

p

p

p

ξ δ−

+

+

+

+

( ) 2 2 2

1 2 3

0 ( )

1 1 0

(

)

(|

|

|

|

|

| )

2

,

t s

t s

e

p

p

p

ds

e

dp p ds

ξ δ

ξ δ

ξ δ

=

+

+

+

( )

2 2 0

2

t

e

ξ δ− s

dp p ds

,

+

( )

3 3 0

2

t

e

ξ δ− s

dp p ds

,

+

( ) 2 2 2

1 2 2 2 3 2

0

(

) .

t s

e

ξ δ−

f

f

f

ds

+

‖ ‖ ‖ ‖ ‖ ‖

+

+

(3)

then

( ) 2 2 2

1 2 3

2 2 2

10 20 30

(|

|

|

|

|

| )

(|

|

|

|

|

| )

t

e

E

p

p

p

E

p

p

p

ξ δ−

+

+

=

+

+

( ) 2 2 2

1 2 3

0

(

ξ δ

)

t

e

ξ δ− s

E

(|

p

|

|

p

|

|

p

| )

ds

+ −

+

+

( )

1 1 0

2

E

t

e

ξ δ− s

dp p ds

,

+

( )

2 2 0

2

E

t

e

ξ δ− s

dp p ds

,

+

( )

3 3 0

2

E

t

e

ξ δ− s

dp p ds

,

+

( ) 2 2 2

1 2 2 2 3 2

0

(

) .

t s

E

e

ξ δ−

f

f

f

ds

+

‖ ‖ ‖ ‖ ‖ ‖

+

+

(4)

( )

1 1 0

( ) 1

1 0

2

,

2

,

t s t s

E

e

dp p ds

p

E

e

p ds

a

ξ δ ξ δ − −

= −

〈−

( )

1 1 1

0 ( )

1 2 1 1

0

2

( , )

,

2

( , )

( )

,

.

t s

t s

E

e

a s p p ds

E

e

a s P t p p ds

ξ δ ξ δ

µ

λ

− −

(5)

By (H1) − (H6)

( )

1 1 0

2

E

t

e

ξ δ− s

dp p ds

,

0 2 2 2

1 1 10 1 1 1 1

( ) 2 2 2

1 2 3

0

[(

)

2

2

2

]

(|

|

|

|

|

| )

.

t s

M a

A r

B r

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

(6)

( )

2 2 0

2

E

t

e

ξ δ− s

dp p ds

,

0 2 2 2

2 2 20 2 2 3 3

( ) 2 2 2

1 2 3

0

[(

)

2

2

2

]

(|

|

|

|

|

| )

.

t s

M a

A r

B r

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

(7)

( )

3 3 0

2

E

t

e

ξ δ− s

dp p ds

,

0 2 2 2

3 3 30 3 2

( ) 2 2 2

1 2 3

0

[(

)

2

2

]

(|

|

|

|

|

| )

.

t s

M a

A r

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

(8)

By Lipschitz(H7),

2 2

2

|

| ,

1, 2, 3.

i i i

f

K

p

i

=

‖ ‖

Then

( ) 2 2 2

1 2 3

2 2 2

10 20 30

(|

|

|

|

|

| )

(|

|

|

|

|

| )

t

e

E

p

p

p

E

p

p

p

ξ δ−

+

+

+

+

( ) 2 2 2

1 2 3

0

(

)

t s

(|

|

|

|

|

| )

e

ξ δ

E p

p

p

ds

ξ δ

+ −

+

+

0 2 2 2

1 1 10 1 1 1 1 1

( ) 2 2 2

1 2 3

0

[(

)

2

2

2

]

(|

|

|

|

|

| )

t s

M a

A r

B r

K

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

+

+

0 2 2 2

2 2 20 2 2 3 3 2

( ) 2 2 2

1 2 3

0

[(

)

2

2

2

]

(|

|

|

|

|

| )

t s

M a

A r

B r

K

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

+

+

0 2 2 2

3 3 30 3 2 3

( ) 2 2 2

1 2 3

0

[(

)

2

2

]

(|

|

|

|

|

| )

t s

M a

A r

K

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

+

+

(9)

2 2 2

10 20 30

( ) 2 2 2

1 2 3

0

(|

|

|

|

|

| ) (

)

(|

|

|

|

|

| )

.

t s

E

p

p

p

e

ξ δ

E

p

p

p

ds

ξ δ τ

+

+

+

− −

+

+

where,

0 2 2 2 0

2

Br

2

(

)

M a

2

AR

K

,

τ

=

+

µ

β

+

,

i

K

=

maxK

A

=

maxA

i

,

R

=

maxr

i

,

0

max

i0

,

β

=

β

µ

0

=

min

µ

i0

,

B

=

minB

i

,

,

i

r

=

minr

i

=

1, 2, 3.

On applying the by Gronwall’s lemma ,we then have the following inequality:

2 2 2

1 2 3

(|

|

|

|

|

| )

t

,

0

E

p

+

p

+

p

Ce

−τ

∀ ≥

t

The proof is proved.

We shall assume the following generalized coercivity condition (H):

There exist constants

α

>

0,

ξ

>

0,

λ

R

,

and a

nonnegative continuous function

γ

( ),

t

t

R

+

,

such that

2 2 2

2

( ,

i

)

i

|

i

|

( )

t

,

f t p

≤ −

α

p

+

λ

p

+

γ

t e

−ξ

‖ ‖

2 3 3

(0, ;

(0,

;

))

(

;

),

. . ,

i T

p

C

T L

a R

+

L Q R

a e t

∀ ∈

where,for arbitrar

δ

>

0,

γ

( )

t

satisfies

( )

t

o e

(

δt

),

γ

=

as

, . ., lim ( ) /

t

0,

t

t

i e

γ

t

e

δ

→∞

→ ∞

=

As follows we give another theorem of this paper.

Theorem 3.2 Assume the preceding hypotheses and (H) hold, if

(

p p p

1

,

2

,

3

)

is a global strong solution to Eq(2). there exist contants

ν

>

0,

C

>

0

such that

2 2 2

1 2 3

(|

|

|

|

|

| )

t

,

0

E

p

+

p

+

p

Ce

−ν

∀ ≥

t

(10)

Proof. We can choose

δ

>

0

small enough such that

ξ δ

− >

0

Then, by

Ito

ˆ

formula , we have

( ) 2 2 2

1 2 3

(|

|

|

|

|

| )

t

(5)

ISSN: 1992-8645 www.jatit.org E-ISSN: 1817-3195

2 2 2

10 20 30

(|

|

|

|

|

| )

E

p

p

p

=

+

+

( ) 2 2 2

1 2 3

0

(

ξ δ

)

t

e

ξ δ− s

E

(|

p

|

|

p

|

|

p

| )

ds

+ −

+

+

( ) ( )

1 1 2 2

0 0

2

E

t

e

ξ δ− s

dp p ds

,

2

E

t

e

ξ δ− s

dp p ds

,

+

〉 +

( )

3 3 0

( ) 2 2 2

1 2 2 2 3 2 0

2

,

(

)

.

t s t s

E

e

dp p ds

E

e

f

f

f

ds

ξ δ ξ δ − −

+

+

+

+

‖ ‖ ‖ ‖ ‖ ‖

(11)

By (H1) − (H6)

( ) 2 2 2

1 2 3

(|

|

|

|

|

| )

t

e

ξ δ−

E

p

+

p

+

p

2 2 2

10 20 30

(|

|

|

|

|

| )

E

p

p

p

+

+

( ) 2 2 2

1 2 3

0

(

ξ δ

)

t

e

ξ δ− s

E

(|

p

|

|

p

|

|

p

| )

ds

+ −

+

+

0 2 2 2

1 1 10 1 1 1 1

[(

β

)

M a

+

2

µ

2

A r

2

B r

]

+

+

( ) 2 2 2

1 2 3

0

(|

|

|

|

|

| )

t s

e

ξ δ−

E

p

+

p

+

p

ds

0 2 2 2

2 2 20 2 2 3 3

( ) 2 2 2

1 2 3

0

[(

)

2

2

2

]

(|

|

|

|

|

| )

t s

M a

A r

B r

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

+

0 2 2 2

3 3 30 3 2

( ) 2 2 2

1 2 3

0

[(

)

2

2

]

(|

|

|

|

|

| )

t s

M a

A r

e

ξ δ

E

p

p

p

ds

β

+

µ

+

+

+

+

( ) 2 2 2

1 2 2 2 3 2

0

(

)

t s

E

e

ξ δ−

f

f

f

ds

+

‖ ‖ ‖ ‖ ‖ ‖

+

+

2 2 2

10 20 30

(|

|

|

|

|

| )

E

p

p

p

+

+

0 2 2 2

0

(

ξ δ

(

β

)

M a

+

2

AR

2

Br

2

µ

)

+ − +

+

( ) 2 2 2

1 2 3

0

(|

|

|

|

|

| )

t s

e

ξ δ−

E

p

+

p

+

p

ds

( ) 2 2 2

1 2 2 2 3 2

0

(

) .

t s

E

e

ξ δ−

f

f

f

ds

+

‖ ‖ ‖ ‖ ‖ ‖

+

+

(12)

0 0

,

i

,

i

,

i

where A

=

maxA R

=

maxr

β

=

max

β

,

0 0

,

,

i i

M

=

maxM

µ

=

min

µ

B

=

minB

i ,

i

r

=

minr

, i=1,2,3.

By (H)

( ) 2 2 2

1 2 3

(|

|

|

|

|

| )

t

e

ξ δ−

E

p

+

p

+

p

2 2 2

10 20 30

(|

|

|

|

|

| )

E

p

p

p

+

+

0 2 2 2

0

(

ξ δ

(

β

)

M a

+

2

AR

2

Br

2

µ

)

+ − +

+

( ) 2 2 2

1 2 3

0

(|

|

|

|

|

| )

t s

e

ξ δ−

E

p

+

p

+

p

ds

( ) 2 2 2

1 2 2 2 3 2

0

(

)

t s

e

ξ δ

E

p

p

p

ds

α

‖ ‖ ‖ ‖ ‖ ‖

+

+

( ) 2 2 2

1 2 3

0

(|

|

|

|

|

| )

t s

e

ξ δ

E

p

p

p

d

λ

+

+

+

0

3

t

γ

( )

s e

−δs

ds

+

2 2 2

10 20 30

(|

|

|

|

|

| )

E

p

p

p

+

+

( ) 2 2 2 1 2 3 0

( ) t s (| | | | | | )

eξ δ E p p p ds

ξ δ ν

+ − −

+ +

0

3

t

γ

( )

s e

−δs

ds

.

+

(13)

Where

ν α

=

/

m

2

− +

λ

2

Br

+

2

µ

0

0 2 2 2

(

β

)

M a

+

2

AR

,

If

ξ ν

− ≤

0

, it follows immediately

( ) 2 2 2

1 2 3

2 2 2

10 20 30

0

(| | | | | | )

(| | | | | | ) 3 ( ) .

t

t

s

e E p p p

E p p p s e ds

ξ δ δ

γ

− − + +

≤ + + +

(14

)

which means that there exists a positive constant

( )

k

=

k

δ

such that

2 2 2

1 2 3

2 2 2 ( )

10 20 30

(|

|

|

|

|

| )

( (|

|

|

|

|

| )

( )

t

.

E p

p

p

E p

p

p

k

δ

e

− −ξ δ

+

+

+

+

+

(15

)

On the other hand , if

ξ ν

− >

0

, we can choose

0

δ

>

small enough such that

ξ δ ν

− − >

0.

Then , from (13) and by Gronwall’s lemma can obtain

( ) 2 2 2

1 2 3

2 2 2

10 20 30

( )

0

(|

|

|

|

|

| )

( (|

|

|

|

|

| )

3

( )

)

.

t

t

s t

e

E

p

p

p

E

p

p

p

s e

ds e

ξ δ

δ ξ δ ν

γ

− − − −

+

+

+

+

+

and, once again, there exists a positive constant

( )

0

k

δ

>

such that

2 2 2

1 2 3

2 2 2

10 20 30

(|

|

|

|

|

| )

( (|

|

|

|

|

| )

(

)

t

.

E

p

p

p

E

p

p

p

k

δ

e

−ν

+

+

+

+

+

The proof of this Theorem3.2 is complete.

(6)

species with age structure by Theorem3.1and Theorem3.2.

4. CONCLUSION

In this paper, we discussed the exponential stability of strong solution for Eq(2). As the exponential stability of strong solution for a stochastic disturbance predator-prey system of three species with age-structure is reached out under the condition of requirements from (H1) to (H7) and (H) being satisfied, we plan to weaken the conditions in the future to substitute local Lipschitz to global Lipschitz and make a further proof of exponential stability of Eq(2).

REFRENCES:

[1] C.Rorres and W. Fair , “Optimal age specific harvesting policy for continuous-time population model ,” in: T.A.Burton(ED.), Modeling and Differential Equations in Biology,Dekker, New York,1980.

[2] W.L. Chan and B.Z. Guo, “Optimal birth control of population dynamics,” J. Math. Anal. Appl,vol. 144, 1989, pp. 532-552.

[3] W.L. Chan and B.Z. Guo, “Optimal birth control of population dynamics. Part 2.Problems with free final time,phase constraints,and mini-max costs,” J.Math.Anal.Appl,vol.146, 1990, pp. 523-539.

[4] S. Anita, ”Optimal harvesting for a nonlinear age-dependent population

dynamics,” J.Math.Anal.Appl, vol. 226,1998, pp. 6-22.

[5] B.Ainseba and M.Langlais, “On a population dynamics control problem with age dependent

and spatial structure,” J.Math.Anal.Appl,vol.248,2000, pp. 455-474.

[6] V. Barbu, M. Iannelli, M. Martcheva, On the controllability of the LotkaCMckendrick model of population dynamics, J. Math. Anal. Appl. Vol. 253, 2001, pp. 142C165.

[7] V. Barbu, M. Iannelli, “Optimal control of population dynamics”, J. Optim. Theo. Appl. Vol. 102, 1999, pp. 1C14.

[8] B.Z. Guo, G. Zhu, “Control Theory of Population Distributional Parameter Systems”, Press of Central China University of Science and Technology, Wuhan (in Chinese), 1999. [9] S. Anita, “Analysis and Control of

Age-Dependent Population Dynamics”, Kluwer Academic Publishers, Dordrecht, 2000.

[10] G.F.Webb,“Theory of Nonlinear Age-dependent Population Dynamics”, Marcel Dekker, NY, 1985.

[11] Zhixue Luo, Ze-Rong He and Wan-Tong Li, “Optimal birth control for Predator Cprey system of three species with age-structure”,Applied Mathematics and Computation . Vol. 155, 2004, pp. 665C685. [12] Tan DJ, Zhang SW, “Permanence in A Delayed

Stage-Structured Three Species Predator-Prey System with Impulsive Perturbations on Predators”, PROCEEDINGS OF THE 7TH CONFERENCE ON BIOLOGICAL DYNAMIC SYSTEM AND STABILITY OF DIFFERENTIAL EQUATION, VOLS I AND II. 2010, pp. 272-278.

[13] Liu ZJ, “Existence of Periodic Solutions for a Delayed Ratio-Dependent Three-Species Predator-Prey Diffusion System on Time Scalese”,ADVANCES IN DIFFERENCE EQUATIONS. 2009, 141589.

[14] Xu R, Chen LS, “Persistence and global stability for a three-species ratio-dependent predator-prey system with time delays in two-patch environments”, ACTA MATHEMATICA SCIENTIA. Vol. 22, No. 4, OCT 2002, pp.533-541.

[15] Zhang YP, Sun JT. ”Persistence in a three species Lotka-Volterra non-periodic Predator-Prey system”, APPLIED MATHEMATICS AND MECHANICS-ENGLISH EDITION. Vol. 21, No. 8, AUG 2000, pp. 879-884.

References

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