Species Author AFS Common Name Change Notes Family Acroloxidae 1 Genus 1 Species. Family Lymnaeidae 14 Genera 62 Species

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The 2021 FMCS checklist of 729 freshwater gastropods from Canada and the United States. Column headings are taxon (binomen) and species author(s), and AFS common names (Classification Uncertain denotes uncertain taxonomic status under common names), Family names are followed by the number of genera and species (or monotypic). Species names denoted in

BLUE LETTERS signify petition changes completed in 2019. Species names denoted in RED LETTERS signify petitioned

changes completed in 2021. The goal of the FMCS Names Subcommittee is to improve science communication, provide up-to- date authoritative checklists and promote freshwater mollusk systematics research. Stakeholders are under no obligation to follow the taxonomic opinions expressed in this list.

Species Author AFS Common Name Change Notes

Family Acroloxidae 1 Genus 1 Species

Acroloxus coloradensis (Henderson, 1930) Rocky Mountain Capshell

Family Lymnaeidae 14 Genera 62 Species

Acella haldemani (Binney, 1867) Spindle Lymnaea

Bulimnaea megasoma (Say, 1824) Mammoth Lymnaea

Erinna aulacospira (Ancey, 1899) Hawaiian Bugle

Erinna newcombi H. Adams & A. Adams, 1855 Newcomb's Bugle

Fisherola nuttalli (Haldeman, 1841) Shortface Lanx

Galba alberta F.C. Baker, 1919 Alberta Fossaria

Galba bulimoides (I. Lea, 1841) Prairie Fossaria

Galba cockerelli (Pilsbry & Ferriss, 1906) Classification Uncertain

Galba cubensis (L. Pfeiffer, 1839) Carib Fossaria

Galba cyclostoma (Walker, 1908) Bugle Fossaria

Galba dalli (F.C. Baker, 1907) Dusky Fossaria

Galba exigua (I. Lea, 1841) Graceful Fossaria

Galba galbana (Say, 1825) Boreal Fossaria

Galba humilis (Say, 1822) Marsh Fossaria

Galba modicella (Say, 1825) Rock Fossaria

Galba obrussa (Say, 1825) Golden Fossaria

Galba parva (I. Lea, 1841) Pygmy Fossaria

Galba peninsulae (B. Walker, 1908) Classification Uncertain

Galba perplexa (Baker & Henderson, 1929) Classification Uncertain

Galba perpolita (Dall, 1905) Glossy Fossaria

Galba rustica (I. Lea, 1841) Rusty Fossaria

Galba sonomaensis (Hemphill, 1906) Sonoma Fossaria

Galba tazewelliana (Wolf, 1870) Tazwell Fossaria

Galba techella (Haldeman, 1867) Classification Uncertain

Galba truncatula (O. F. Müller, 1774) Attenuate Fossaria

Galba vancouverensis F.C. Baker, 1939 Classification Uncertain

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Hinkleyia caperata (Say, 1829) Wrinkled Marshsnail

Placed into Hinkleyia Jan. 2020 by M. Vinarski.

FMCS petition G-2021-01.

Hinkleyia pilsbryi (Hemphill, 1890) Fish Springs Marshsnail

Placed into Hinkleyia Jan. 2020 by M. Vinarski.

FMCS petition G-2021-01.

Idaholanx fresti Clark, Campbell & Lydeard, 2017 Banbury Springs Limpet

Formal description of Banbury Springs Limpet, Campbell et al. ZooKeys 2017 663:107-132. FMCS petition G-2019-5.

Ladislavella atkaensis (Dall, 1884) Frigid Lymnaea Placed in Ladislavella FMCS petition G-2021-02.

Ladislavilla eloides (Say, 1821)

Taxon mistakenly excluded from Johnson et al.

2013. Placed in Ladislavella FMCS petition G-2021- 02.

Ladislavella apicina (I. Lea, 1838) Abbreviate Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella arctica (I. Lea, 1864) Arctic Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella bonnevillensis (Call, 1884) Fat-Whorled Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella catascopium (Say, 1817) Woodland Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella contracta (Currier in DeCamp, 1881) Deepwater Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella elodes (Say, 1821) Marsh Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella elrodi (F.C. Baker & Henderson, 1933) Flathead Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella elrodiana (F.C. Baker, 1935) Longmouth Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella emarginata (Say, 1821) St. Lawrence Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella exilis (I. Lea, 1834) Flat-Whorled Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella gabbi (Tryon, 1865) Striate Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella hinkleyi (F.C. Baker, 1906) Rustic Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella idahoensis (Henderson, 1931) Shortspire Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella kennicotti (F.C. Baker, 1933) Western Arctic Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella mighelsi (Binney, 1865) Bigmouth Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella neopalustris (F.C. Baker, 1911) Piedmont Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella oronoensis (F.C. Baker, 1904) Obese Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella petoskeyesis (B. Walker, 1908) Petosky Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella traskii (Tryon, 1863) Widelip Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella utahensis (Call, 1884) Thickshell Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Ladislavella woodruffi (F.C. Baker, 1901) Coldwater Pondsnail Placed in Ladislavella FMCS petition G-2021-02.

Lanx alta (Tryon, 1865) Highcap Lanx

Lanx klamathensis Hannibal, 1912 Scale Lanx

Lanx patelloides (Lea, 1856) Kneecap Lanx

Lanx subrotunda (Tryon, 1865) Rotund Lanx

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Lymnaea stagnalis (Linnaeus, 1758) Swamp Lymnaea

Taxon erroneously excluded from Johnson et al.

2013. Accidental deletion.

Pseudisdora producta (Mighels, 1845) Classification Uncertain Placed in Pseudisidora FMCS peptition G-2021-03

Pseudisidora rubella (Lea, 1841) Aloha Lymnea Placed in Pseudisidora FMCS peptition G-2021-03

Pseudosuccinea columella (Say, 1817) Mimic Lymnaea

Stagnicola walkeriana (F.C. Baker, 1926) Calabash Pondsnail

Walterigalba montanensis (F.C. Baker, 1913) Mountain Marshsnail Placed in Walterigalbia FMCS petition G-2021-04.

Family Physidae 3 Genera 45 Species

Aplexa elongata (Say, 1821) Lance Aplexa

Aplexa microstriata (Chamberlin & Berry, 1930) Fish Lake Aplexa

Physa jennessi Dall, 1919 Obtuse Physa

Physa chippuvarum (Taylor, 2003) Chippewa Physa

Molluscabase and Lydeard & Cummings 2019 and reassignment of Laurentiphysa to Physa, FMCS Petition G-2019-6.

Physa megalochlamys Taylor, 1988 Cloaked Physa

Physa sibirica (Westerlund, 1876) Frigid Physa

Physa skinneri Taylor, 1954 Glass Physa

Physa vernalis (Taylor & Jokinen, 1984) Vernal Physa

Physella acuta (Draparnaud, 1805) Wandering Physa

Returned to list from analyses by Lydeard, Campbell, and Gotz, 2016. Malacologia. 59(2):347- 350. FMCS petition G-2019-12. Physella mexicata (Philippi in Kuster, 1841) and Physella virgata Gould 1855 and Physella winnipegensis were

synonymized with Physella acuta, Petition Gastropod 2021-8.

Physella ancillaria (Say, 1825) Pumpkin Physa

Physella ashmuni (Taylor, 2003) San Rafael Physa

Archiphysai Taylor, 2003 synonomized with Physella by Wethington and Lydeard 2007, as revised by Vinarski 2018. FMCS Petition G-2019-01.

Petition tabled but adopted in 2021.

Physella bermudezi (Aguayo, 1935) Lowdome Physa

Physella bottimeri (Clench, 1924) Comanche Physa

Physella boucardi (Crosse & Fischer, 1881) Desert Physa

Physella carolinae Wethington, Dillon, Wise, 2009 Carolina Physa

Physella columbiana (Hemphill, 1890) Rotund Physa

Physella conoidea (Fischer & Crosse, 1886) Texas Physa

Physella cooperi (Tryon, 1865) Olive Physa

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Physella costata (Newcomb, 1861) Ornate Physa

Physella cubensis Pfeiffer, 1839 Carib Physa

Physella globosa (Haldeman, 1842) Globose Physa

Physella gyrina (Say, 1821) Tadpole Physa

Physella hemphilli Taylor, 2003 Idaho Physa

Physella hendersoni (Clench, 1925) Bayou Physa

Physella heterostropha (Say, 1817) Pewter Physa

Physella hordacea (I. Lea, 1864) Grain Physa

Physella humerosa (Gould, 1855) Corkscrew Physa

Physella integra (Haldeman, 1841) Ashy Physa

Physella johnsoni (Clench, 1926) Banff Springs Physa

Physella lordi (Baird, 1863) Twisted Physa

Physella magnalacustris (Walker, 1901) Great Lakes Physa

Physella natricina (Taylor, 1988) Snake River Physa Placed in Physella Gates et al., 2013.

Physella osculans (Haldeman, 1841) Cayuse Physa

Physella parkeri (Currier in DeCamp, 1881) Broadshoulder Physa

Physella pomilia (Conrad, 1834) Claiborne Physa

Physella propinqua (Tryon, 1865) Rocky Mountain Physa

Physella spelunca (Turner & Clench, 1974) Cave Physa

Physella squalida (Morelet, 1851) Squalid Physa

Physella sonomae (Taylor, 2003) Sonoma Physa

ArchiphysaTaylor, 2003 synonomized with Physella by Wethington and Lydeard 2007, as revised by Vinarski 2018. FMCS Petition G-2019-01. Petition tabled but adopted in 2021.

Physella traskii (I. Lea, 1864) Sculpted Physa

Physella utahensis (Clench, 1925) Utah Physa

Physella vinosa (Gould, 1847) Banded Physa

Physella virginea (Gould, 1847) Sunset Physa

Physella wrighti Te & Clarke, 1985 Hotwater Physa

Physella zionis (Pilsbry, 1926) Wet-rock Physa

Family Planorbidae 17 Genera 53 Species

Amphigyra alabamensis Pilsbry, 1906 Shoal Sprite

Armiger crista (Linnaeus, 1758) Star Gyro Taxon reviewed Petition Gastropod 2021-10

Biomphalaria havanensis (Pfeiffer, 1839) Ghost Ramshorn

Dilatata brogniartiana (Lea, 1842) Disc Sprite

Micomenetus placed in Dilatata. Petition Gastropod 2019-09. Petition tabled but adopted for Dilatata 2021.

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Dilatata dilatata (Gould, 1841) Bugle Sprite Petition Gastropod 2019-09.

Dilatata floridensis (Baker, 1945) Penny Sprite Petition Gastropod 2019-09.

Dilatata sampsoni (Ancey, 1885) Sampson Sprite Petition Gastropod 2019-09.

Ferrissia californica (Rowell, 1863) Fragile Ancylid Ferrissia californica (Rowell, 1863) is the priority name for F. fragilis. Petition Gastropod 2021-09

Ferrissia rivularis (Say, 1817) Creeping Ancylid

Gyraulus circumstriatus (Tryon, 1866) Disc Gyro

Gyraulus crista (Linnaeus, 1758) Star Gyro

Gyraulus deflectus (Say, 1824) Flexed Gyro

G. hornensis F.C. Baker, 1934 and G. vermicularis (Gould, 1947) are not synonymized with G.

deflectus (Say, 1824). Petition Gastropod 2021-11

Gyraulus hornensis Baker, 1934 Tuba Gyro

Gyraulus parvus (Say, 1817) Ash Gyro

Gyraulus vermicularis (Gould, 1847) Pacific Coast Gyro

Hebetancylus excentricus (Morelet, 1851) Eccentric Ancylid

Helisoma anceps (Menke, 1830) Two-ridge Ramshorn

Helisoma minus (Cooper, 1870) Classification Uncertain

Helisoma newberryi (Lea, 1858) Great Basin Ramshorn

Laevapex fuscus (Adams, 1840) Dusky Ancylid

Menetus opercularis (Gould, 1847) Button Sprite

Neoplanorbis carinatus Walker, 1908 Carinate Flat-top Snail

Neoplanorbis smithi Walker, 1908 Angled Flat-top Snail

Neoplanorbis tantillus Pilsbry, 1906 Little Flat-top Snail

Neoplanorbis umbilicatus Walker, 1908 Umbilicate Flat-top Snail

Pecosorbis kansasensis (Berry, 1966) New Mexico Ramshorn

Planorbella ammon (Gould, 1855) Jupiter Ramshorn

Planorbella binneyi (Tryon, 1867) Coarse Ramshorn

Planorbella campanulata (Say, 1821) Bellmouth Ramshorn

Planorbella columbiensis (Baker, 1945) Caribou Ramshorn

Planorbella corpulenta (Say, 1824) Corpulent Ramshorn

Planorbella duryi (Wetherby, 1879) Seminole Ramshorn

Planorbella magnifica (Pilsbry, 1903) Magnificent Ramshorn

Planorbella multivolvis (Case, 1847) Acorn Ramshorn

Planorbella occidentalis (Cooper, 1870) Fine-lined Ramshorn

Planorbella oregonensis (Tryon, 1865) Lamb Ramshorn

Planorbella pilsbryi (Baker, 1926) File Ramshorn

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Planorbella scalaris (Jay, 1839) Mesa Ramshorn

Planorbella subcrenata (Carpenter, 1857) Rough Ramshorn

Planorbella tenuis (Dunker, 1850) Mexican Ramshorn

Planorbella traski (Lea, 1856) Keeled Ramshorn

Planorbella trivolvis (Say, 1817) Marsh Ramshorn

Planorbella truncata (Miles, 1861) Druid Ramshorn

Planorbula armigera (Say, 1821) Thicklip Ramshorn

Planorbula campestris (Dawson, 1875) Meadow Ramshorn

Promenetus exacuous (Say, 1821) Sharp Sprite

Promenetus umbilicatellus (Cockerell, 1887) Umbilicate Sprite

Rhodacmea cahawbensis (Walker, 1917) Cahaba Ancylid

Rhodacmea elatior (J. G. Anthony, 1855) Domed Ancylid

Rhodacmea filosa (Conrad, 1834) Wicker Ancylid

Rhodacmea hinkleyi (Walker, 1908) Knobby Ancylid

Vorticifex effusa (I. Lea, 1856) Artemesian Ramshorn

Vorticifex solida (Dall, 1870) Classification Uncertain

Family Neritidae 1 Genus 5 Species

Neritina cariosa (Wood, 1828) Pip'wai

Neritina granosa Sowerby, 1825 Hihiwai

Neritina vespertina Sowerby, 1849 Hapawai

Neritina clenchi Russel, 1940 Classification Uncertain FMCS petition G-2021-12 to place gulf coast Neritina into Vitta is tabled until 2023

Neritina usnea (Roding, 1798) Olive Nerite FMCS petition G-2021-12 to place gulf coast

Neritina into Vitta is tabled until 2023.

Family Viviparidae 4 Genera 21 Species

Callinina georgiana (Lea, 1834) Banded Mysterysnail New combination Stelbrink et al. 2020. Petition

Gastropod 2021-13.

Callinina goodrichi (Archer, 1933) Globose Mysterysnail

Callinina intertexta (Say, 1829) Rotund Mysterysnail

Callinina limi (Pilsbry, 1918) Ochlocknee Mysterysnail

Callinina subpurpura (Say, 1829) Olive Mysterysnail

Campeloma brevispirum Baker, 1928 Classification Uncertain

Campeloma crassulum Rafinesque, 1819 Ponderous Campeloma

Campeloma decampi (Binney, 1865) Slender Campeloma

Campeloma decisum (Say, 1817) Pointed Campeloma

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Campeloma floridense Call, 1886 Purple-throat Campeloma

Campeloma geniculum (Conrad, 1834) Ovate Campeloma

Campeloma limum (Anthony, 1860) File Campeloma

Campeloma milesi (Lea, 1863) Classification Uncertain

Campeloma parthenum Vail, 1979 Maiden Campeloma

Campeloma regulare (Lea, 1841) Cylinder Campeloma

Campeloma rufum (Haldeman, 1841) Classification Uncertain

Lioplax cyclostomaformis (Lea, 1841) Cylindrical Lioplax

Lioplax pilsbryi Walker, 1905 Choctaw Lioplax

Lioplax subcarinata (Say, 1817) Rigid Lioplax

Lioplax sulculosa (Menke, 1827) Furrowed Lioplax

Tulotoma magnifica (Conrad, 1834) Tulotoma

Family Ampullariidae 1 Genus 1 Species

Pomacea paludosa (Say, 1829) Florida Applesnail

Family Assimineidae 2 Genera 2 Species

Angustassiminea infima (Berry, 1947) Badwater Snail

Assiminea infima Berry, 1947 was placed in Angustassiminea Habe, 1943. Petition Gastropod 2021-14.

Assiminea pecos Taylor, 1987 Pecos Assiminea

Family Amnicolidae 4 Genera 20 Species

Amnicola cora Hubricht, 1979 Foushee Cavesnail

Amnicola dalli (Pilsbry & Beecher, 1892) Peninsula Amnicola

Amnicola decisus Haldeman, 1845 Classification Uncertain

Amnicola limosus (Say, 1817) Mud Amnicola

Amnicola rhombostomus Thompson, 1968 Squaremouth Amnicola

Amnicola stygius Hubricht, 1971 Stygian Amnicola

Colligyrus convexus Hershler, Frest, Liu, & Johannes,

2003 Canary Duskysnail

Colligyrus depressus Hershler, 1999 Harney Basin Duskysnail

Colligyrus greggi (Pilsbry, 1935) Rocky Mountain Duskysnail

Dasyscias franzi Thompson & Hershler, 1991 Shaggy Ghostsnail

Lyogyrus bakerianus (Pilsbry, 1917) Baker's Springsnail

Lyrogyrus browni (Carpenter, 1872) Slender Duskysnail

Lyogyrus granum (Say, 1822) Squat Duskysnail

Lyogyrus latus Thompson & Hershler, 1991 Cobble Sprite

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Lyogyrus pilsbryi (Walker, 1906) Lake Duskysnail

Lyogyrus pupoideus (Gould, 1841) Pupa Duskysnail

Lyogyrus retromargo (Thompson, 1968) Indented Duskysnail

Lyogyrus sp. Liu et al., 2016 Masked Duskysnail Recent review Liu et al, 2016 - JMS. Diagnosis

pending.

Lyogyrus walkeri (Pilsbry, 1898) Canadian Duskysnail

Family Cochliopidae 19 Genera 63 Species

Antrobia culveri Hubricht, 1971 Tumbling Creek Cavesnail

Antrorbis breweri Hershler & Thompson, 1990 Manitou Cavesnail

Reassigned from Lithoglyphidae. Petition Gastropod 2021-16.

Antrorbis tennesseensis Perez, Shoobs, Gladstone, &

Niemiller, 2019 Tennessee Cavesnail

Antrorbis tennesseensis Perez et al. 2019.

Recognized Petition Gastropod 2021-16.

Antroselates spiralis Hubricht,1963 Shaggy Cavesnail

Aphaostracon asthenes F. G. Thompson, 1968 Blue Spring Hydrobe

Aphaostracon chalarogyrum F. G. Thompson, 1968 Freemouth Hydrobe

Aphaostracon hypohyalinum F. G. Thompson, 1968 Suwanee Hydrobe

Aphaostracon monas (Pilsbry, 1899) Wekiwa Hydrobe

Aphaostracon pachynotum F. G. Thompson, 1968 Thick-shelled Hydrobe

Aphaostracon pycnum F. G. Thompson, 1968 Dense Hydrobe

Aphaostracon rhadinum F. G. Thompson, 1968 Slough Hydrobe

Aphaostracon theiocrenetum F. G. Thompson, 1968 Clifton Spring Hydrobe Aphaostracon xynoelictum F. G. Thompson, 1968 Fenney Spring Hydrobe Balconorbis uvaldensis Hershler & Longley, 1986 Balcones Ghostsnail Cochliopina riograndensis Pilsbry & Ferriss, 1906 Spiral Pebblesnail

Eremopyrgus eganensis Hershler, 1999 Steptoe Hydrobe

Holsingeria unthanksensis Hershler, 1988 Thankless Ghostsnail

Holsingeria unthanksensis Hershler, 1988 was placed in the Cochliopidae. Petition Gastropod 2021-15.

Ipnobius robustus (Hershler, 1989) Robust Tryonia

Juturnia brunei (Taylor, 1987) Brune's Springsnail

Juturnia kosteri (Taylor, 1987) Koster Springsnail

Juturnia tularosae Hershler, Liu, & Stockwell, 2002 Tularosa Springsnail

Littoridinops monroensis (Frauenfeld, 1863) Cockscomb Hydrobe

Littoridinops palustris F. G. Thompson, 1968 Bantam Hydrobe

Littoridinops tenuipes (Couper, 1844) Henscomb Hydrobe

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Phreatoceras taylori (Hershler & Longley, 1986) Nymph Trumpet Phreatoceras moved to Cochliopidae. Petition Gastropod 2019-10.

Phreatodrobia conica Hershler & Longley, 1986 Hueco Cavesnail

Phreatodrobia moved to Cochliopidae FMCS Petition G-2019-10. Taxon erroneously omitted from previous list and was recognized. Petition

Gastropod 2019-11.

Phreatodrobia coronae Hershler & Longley, 1987 Crowned Cavesnail

Phreatodrobia moved to Cochliopidae. Petition Gastropod 2019-10.

Phreatodrobia imitata Hershler & Longley, 1986 Mimic Cavesnail

Phreatodrobia micra (Pilsbry & Ferriss, 1906) Flattened Cavesnail Phreatodrobia nugax (Pilsbry & Ferriss, 1906) Domed Cavesnail

Phreatodrobia plana Hershler & Longley, 1986 Disc Cavesnail

Phreatodrobia punctata Hershler & Longley, 1986 High-hat Cavesnail Phreatodrobia rotunda Hershler & Longley, 1986 Beaked Cavesnail

Phreatodrobia spica Perez & Alvear, 2020 Spiky Cavesnail Phreatodrobia spica Perez & Alvear, 2020 recognized Petition Gastropod 2021-17.

Pseudotryonia adamantina (Taylor, 1987) Diamond Tryonia

Pseudotryonia alamosae (Taylor, 1987) Caliente Tryonia

Pseudotryonia brevissima (Pilsbry, 1890) Regal Hydrobe

Pseudotryonia grahamae F. G. Thompson, 1968 Salt Spring Hydrobe

Pyrgophorus platyrachis F. G. Thompson, 1968 Serrate Crownsnail

Pyrgophorus spinosus (Call & Pilsbry, 1886) Spiny Crownsnail

Spurwinkia salsa (Pilsbry, 1905) Saltmarsh Hydrobe

Stygopyrgus bartonensis Hershler & Longley, 1986 Barton Cavesnail

Texapyrgus longleyi Thompson & Hershler, 1991 Striated Hydrobe Texapyrgus was placed in the Cochliopidae. Petition Gastropod 2019-10.

Tryonia aequicostata (Pilsbry, 1890) Smooth-ribbed Hydrobe

Tryonia angulata Hershler & Sada, 1987 Sportingoods Tryonia

Tryonia cheatumi (Pilsbry, 1935) Phantom Tryonia

Tryonia circumstriata (Leonard & Ho, 1960) Gonzales Springsnail

Tryonia clathrata Stimpson, 1865 Grated Tryonia

Tryonia diaboli (Pilsbry & Ferriss, 1906) Devil Tryonia

Tryonia elata Hershler & Sada, 1987 Point of Rocks Tryonia

Tryonia ericae Hershler & Sada, 1987 Minute Tryonia

Tryonia gilae Taylor, 1987 Gila Tryonia

Tryonia imitator (Pilsbry, 1899) Mimic Tryonia

Tryonia infernalis Hershler, Liu, & Simpson, 2015 Fire Valley Tryonia Hershler, Liu, and Simpson, 2015. ZooKeys 492:107-122. Petition Gastropod 2019-23.

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Tryonia margae Hershler, 1989 Grapevine Springs Elongate Springsnail

Tryonia metcalfi Hershler, Liu, & Lanye, 2011 Metcalf's Tryonia

Tryonia monitorae Hershler, 1999 Monitor Tryonia

Tryonia oasiensis Hershler, Liu, & Landye, 2011 Carolinae Tryonia

Tryonia porrecta (Mighels, 1845) Desert Tryonia

Tryonia quitobaquitae Hershler, 1988 Quitobaquito Tryonia

Tryonia rowlandsi Hershler, 1989 Grapevine Springs Squat

Tryonia

Tryonia salina Hershler, 1989 Cottonball Marsh Tryonia

Tryonia variegata Hershler & Sada, 1987 Amargosa Tryonia

Family Emmericiidae 1 Genus 10 Species

Fontigens aldrichi (Call & Beecher, 1886) Hoosier Springsnail New family placement for Fontigens spp. Petition Gastropod 2021-18.

Fontigens antroecetes (Hubricht, 1940) Missouri Cavesnail

Fontigens bottimeri (Walker, 1925) Potomac Springsnail

Fontigens cryptica Hubricht, 1963 Hidden Springsnail

Fontigens morrisoni Hershler, Holsinger & Hubricht, 1990 Morrison's Springsnail

Fontigens nickliniana (Lea, 1838) Watercress Snail

Fontigens orolibas Hubricht, 1957 Blue Ridge Springsnail

Fontigens proserpina (Hubricht, 1940) Proserpine Cavesnail

Fontigens tartarea Hubricht, 1963 Organ Cavesnail

Fontigens turritella Hubricht, 1976 Greenbrier Cavesnail

Family Hydrobiidae 11 Genera 180 Species

Birgella subglobosa (Say, 1825) Globe Siltsnail

Cincinnatia integra (Say, 1821) Midland Siltsnail

Floridobia alexander (Thompson, 2000) Alexander Siltsnail

Floridobia floridana (Frauenfeld, 1863) Hyacinth Siltsnail

Floridobia fraterna (Thompson, 1968) Creek Siltsnail

Floridobia helicogyra (Thompson, 1968) Crystal Siltsnail

Floridobia leptospira (Thompson, 2000) Flatwood Siltsnail

Floridobia mica (Thompson, 1968) Ichetucknee Siltsnail

Floridobia monroensis (Dall, 1885) Enterprise Siltsnail

Floridobia parva (Thompson, 1968) Pygmy Siltsnail

Floridobia petrifons (Thompson, 1968) Rock Springs Siltsnail

Floridobia ponderosa (Thompson, 1968) Ponderous Siltsnail

Floridobia porterae (Thompson, 2000) Green Cove Springsnail

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Floridobia vanhyningi (Vanatta, 1934) Seminole Siltsnail

Floridobia wekiwae (Thompson, 1968) Wekiva Siltsnail

Floridobia winkleyi (Pilsbry, 1912) New England Siltsnail

Hoyia sheldoni (Pilsbry, 1890) Storm Ghostsnail

Marstonia agarhecta Thompson, 1969 Ocmulgee Marstonia

Marstonia angulobasis Thompson, 2005 Angled Marstonia

Marstonia arga Thompson, 1977 Ghost Marstonia

Marstonia castor Thompson, 1977 Beaverpond Marstonia

Marstonia comalensis (Pilsbry & Ferriss, 1906) Comal Marstonia

Marstonia gaddisorum Thompson, 2005 Emily's Marstonia

Marstonia halcyon Thompson, 1977 Halcyon Marstonia

Marstonia hershleri (Thompson, 1995) Coosa Pyrg

Marstonia letsoni (Walker, 1901) Gravel Pyrg

Marstonia lustrica (Pilsbry, 1890) Boreal Marstonia

Marstonia ogmorhaphe Thompson, 1977 Royal Marstonia

Marstonia olivacea (Pilsbry, 1895) Olive Marstonia

Marstonia ozarkensis (Hinkley, 1915) Ozark Pyrg

Marstonia pachyta Thompson, 1977 Armored Marstonia

Marstonia scalariformis (Wolf, 1870) Moss Pyrg

Notogillia sathon Thompson, 1969 Satyr Siltsnail

Notogillia wetherbyi (Dall, 1885) Alligator Siltsnail

Probythinella emarginata (Kuster, 1852) Delta Hydrobe

Pyrgulopsis aardahli Hershler, 1989 Benton Valley Springsnail

Pyrgulopsis aloba Hershler, 1998 Duckwater Pyrg

Pyrgulopsis amargosae Hershler, 1989 Amargosa Springsnail

Pyrgulopsis anatina Hershler, 1998 Southern Duckwater Pyrg

Pyrgulopsis anguina Hershler, 1998 Longitudinal Gland Pyrg

Pyrgulopsis archimedis Berry, 1947 Archimedes Pyrg

Pyrgulopsis arizonae (Taylor, 1987) Apache Springsnail

Pyrgulopsis augustae Hershler, 1998 Elongate Cane Spring Pyrg

Pyrgulopsis aurata Hershler, 1998 Pleasant Valley Pyrg

Pyrgulopsis avernalis (Pilsbry, 1935) Moapa Pebblesnail

Pyrgulopsis bacchus Heshler, 1988 Grand Wash Springsnail

Pyrgulopsis basiglans Hershler, 1998 Large Gland Carico Pyrg

Pyrgulopsis bedfordensis Hershler & Gustafson, 2001 Bedford Pyrg

(12)

Pyrgulopsis bernardina (Taylor, 1987) San Bernardino Springsnail

Pyrgulopsis bifurcata Hershler, 1998 Small Gland Carico Pyrg

Pyrgulopsis blainica Hershler, Liu, Gustafson, 2008 Blane Pyrg

Pyrgulopsis breviloba Hershler, 1998 Flat Pyrg

Pyrgulopsis bruesi Hershler & Sada, 2000 Fy Ranch Pyrg

Pyrgulopsis bruneauensis Hershler, 1990 Bruneau Hot Springsnail

Pyrgulopsis bryantwalkeri Hershler, 1994 Cortez Hills Pebblesnail Pyrgulopsis californiensis (Gregg & Taylor, 1965) Languna Mountain Springsnail

Pyrgulopsis carinata Hershler, 1998 Carinate Duckwater Pyrg

Pyrgulopsis carinifera (Pilsbry, 1935) Moapa Valley Pyrg

Pyrgulopsis castaicensis Hershler & Liu, 2010 Middle Canyon Spring Pyrg

Pyrgulopsis chamberlini Hershler, 1998 Smooth Glenwood Pyrg

Pyrgulopsis chupaderae (Taylor, 1987) Chupadera Springsnail

Pyrgulopsis cinerana Hershler, Frest, Liu, & Johannes,

2003 Ash Valley Pyrg

Pyrgulopsis coloradensis Hershler, 1998 Blue Point Pyrg

Pyrgulopsis conica Hershler, 1988 Kingman Springsnail

Pyrgulopsis cruciglans Hershler, 1998 Transverse Gland Pyrg

Pyrgulopsis crystalis Hershler & Sada, 1987 Crystal Springsnail

Pyrgulopsis cybele Hershler & Liu, 2012 Nature Pyrg

Pyrgulopsis davisi (Taylor, 1987) Limpia Creek Springsnail

Pyrgulopsis deaconi Hershler, 1998 Spring Mountains Pyrg

Pyrgulopsis deserta (Pilsbry, 1916) Desert Springsnail

Pyrgulopsis diablensis Hershler, 1995 Diablo Range Pyrg

Pyrgulopsis dixiensis Hershler, 1998 Dixie Valley Pyrg

Pyrgulopsis eremica Hershler, 1995 Smoke Creek Pyrg

Pyrgulopsis erythropoma (Pilsbry, 1899) Ash Meadows Pebblesnail

Pyrgulopsis fairbanksensis Hershler & Sada, 1987 Fairbanks Springsnail Pyrgulopsis falciglans Hershler, Frest, Liu, & Johannes,

2003 Likely Pyrg

Pyrgulopsis fausta Hershler, 1998 Corn Creek Pyrg

Pyrgulopsis fresti Hershler & Liu, 2009 Owyhee Hot Springsnail

Pyrgulopsis fusca Hershler, 1998 Otter Creek Pyrg

Pyrgulopsis gibba Hershler, 1995 Surprise Valley Pyrg

Pyrgulopsis gilae (Taylor, 1987) Gila Springsnail

Pyrgulopsis giuliani Hershler & Pratt, 1990 Southern Sierra Nevada

Springsnail

(13)

Pyrgulopsis glandulosa Hershler, 1988 Verde Rim Springsnail

Pyrgulopsis gracilis Hershler, 1998 Emigrant Pyrg

Pyrgulopsis greggi Hershler, 1995 Kern River Springsnail

Pyrgulopsis hamlinensis Hershler, 1998 Hamlin Valley Pyrg

Pyrgulopsis hovinghi Hershler, 1998 Upper Thousand Spring Pyrg

Pyrgulopsis hualapaiensis Hershler, Liu, & Stevens, 2016 Hualapai Pyrg

Pyrgulopsis hualapaiensis Hershler et al. 2016 recognized. - WNA Naturalist 76(1):72-81. Petition Gastropod 2019-13.

Pyrgulopsis hubbsi Hershler, 1998 Hubbs Pyrg

Pyrgulopsis humboldtensis Hershler, 1998 Humbolt Pyrg

Pyrgulopsis ignota Hershler, Liu, & Lang, 2010 Caroline Springs Pyrg

Pyrgulopsis imperialis Hershler, 1998 Kings River Pyrg

Pyrgulopsis inopinata Hershler, 1998 Carinate Glenwood Pyrg

Pyrgulopsis intermedia (Tryon, 1865) Crooked Creek Springsnail

Pyrgulopsis isolata Hershler & Sada, 1987 Elongate-gland Springsnail

Pyrgulopsis kolobensis (Taylor, 1987) Tocqueville Springsnail

Pyrgulopsis landyei Hershler, 1998 Landyes Pyrg

Pyrgulopsis lasseni Hershler, Frest, Liu, & Johannes,

2003 Willow Creek Pyrg

Pyrgulopsis lata Hershler, 1998 Butterfield Pyrg

Pyrgulopsis lentiglans Hershler, 1998 Critteden Pyrg

Pyrgulopsis leporina Hershler, 1998 Elko Pyrg

Pyrgulopsis licina Hershler, Liu, & Bradford, 2013 Nye County Pyrg

Pyrgulopsis limaria Hershler, 1998 Squat Mud Meadows Pyrg

Pyrgulopsis lindae Hershler, Liu, Babbitt, Keollogg, &

Howard, 2016 San Domingo Pyrg Pyrgulopsis lindae Hershler et al. 2016 Zookeys

601:1-19. Recognized Petition Gastropod 2019-14.

Pyrgulopsis lindahlae Hershler, Liu, Forsythe, Hovingh &

Wheeler, 2017 Lindahl's Pyrg

Pyrgulopsis lindahlae Hershler et al. 2017 JMS 1- 11. Recognized Petiton Gastropod 2019-15.

Pyrgulopsis lockensis Hershler, 1998 Lockes Pyrg

Pyrgulopsis longae Hershler, 1995 Long Valley Pyrg

Pyrgulopsis longiglans Hershler, 1998 Western Lahontan Pyrg

Pyrgulopsis longinqua (Gould, 1855) Salton Sea Springsnail

Pyrgulopsis marcida Hershler, 1998 Hardy Pyrg

Pyrgulopsis marilynae Hershler, Ratcliffe, Liu, Lang, & Hay,

2014 Jordan Spring Pyrg Pygulopsis marilynae Hershler et al. 2017 JMS 1-

11. Recognized Petition Gastropod 2019-16.

Pyrgulopsis merriami (Pilsbry & Beecher, 1892) Pahrangagat Pebblesnail

Pyrgulopsis metcalfi (Taylor, 1987) Naegele Springsnail

Pyrgulopsis micrococcus (Pilsbry, 1893) Oasis Valley Springsnail

(14)

Pyrgulopsis militaris Hershler, 1998 Northern Soldier Meadow Pyrg

Pyrgulopsis millenaria Hershler, 1998 Twentyone Mile Pyrg

Pyrgulopsis milleri Hershler & Liu, 2010 Pierpoint Spring Pyrg

Pyrgulopsis montana Hershler, 1998 Camp Valley Pyrg

Pyrgulopsis montezumensis Hershler, 1988 Montezuma Well Springsnail

Pyrgulopsis morrisoni Hershler, 1988 Page Springsnail

Pyrgulopsis nanus Hershler & Sada, 1987 Distal-gland Springsnail

Pyrgulopsis neomexicana (Pilsbry, 1916) Socorro Springsnail

Pyrgulopsis neritella Hershler, 1998 Neritiform Steptoe Ranch Pyrg

Pyrgulopsis nevadensis (Stearns, 1883) Corded Pyrg

Pyrgulopsis nuwwvi Hershler, Liu, Forsythe, Hovingh &

Wheeler, 2017 Nuwuvi Pyrg

Pyrgulopsis nuwuvi Hershler et al. 2017 JMS 1-11.

Recognized Petiton Gastropod 2019-17.

Pyrgulopsis nonaria Hershler, 1998 Ninemile Pyrg

Pyrgulopsis notidicola Hershler, 1998 Elongate Mud Meadows Pyrg

Pyrgulopsis ojaiensis Hershler, Liu, Babbitt, Keollogg, &

Howard, 2016 Sisar Pyrg

Pyrgulopsis ojalensis Hershler et al. 2016 Zookeys 601:1-19. Recognized Petition Gastropod 2019-18.

Pyrgulopsis orbiculata Hershler, 1998 Sub-globose Steptoe Ranch Pyrg

Pyrgulopsis owensensis Hershler, 1989 Owens Valley Springsnail

Pyrgulopsis owyheensis Hershler & Liu, 2009 Owyhee Upland Pyrg

Pyrgulopsis papillata Hershler, 1998 Big Warm Spring Pyrg

Pyrgulopsis pecosensis (Taylor, 1987) Pecos Springsnail

Pyrgulopsis peculiaris Hershler, 1998 Bifid Duct Pyrg

Pyrgulopsis pellita Hershler, 1998 Antelope Valley Pyrg

Pyrgulopsis perforata Hershler, Liu, & Bradford, 2013 Umbilicate Pyrg

Pyrgulopsis perturbata Hershler, 1989 Fish Slough Springsnail

Pyrgulopsis pictilis Hershler, 1998 Ovate Cain Spring Pyrg

Pyrgulopsis pilsbryana (Bailey & Bailey, 1952) Bear Lake Springsnail

Pyrgulopsis pinetorum (Taylor, 1987) Pine Grove Pyrg

Pyrgulopsis pinetorum was elevated from synonymy with Pyrgulopsis kolobensis Hershler et al. 2017 JMS 1-11. Petition Gastropod 2019-19.

Pyrgulopsis pisteri Hershler & Sada, 1987 Median-gland Springsnail

Pyrgulopsis planulata Hershler, 1998 Flat-topped Steptoe Pyrg

Pyrgulopsis plicata Hershler, 1998 Black Canyon Pyrg

Pyrgulopsis robusta (Walker, 1908) Jackson Lake Springsnail

Pyrgulopsis roswellensis (Taylor, 1987) Roswell Springsnail

Pyrgulopsis ruinosa Hershler, 1998 Fish Lake Pyrg

(15)

Pyrgulopsis rupinicola Hershler, Frest, Liu, & Johannes,

2003 Sucker Spring Pyrg

Pyrgulopsis sadai Hershler, 1998 Sada's Pyrg

Pyrgulopsis sanchezi Hershler, Liu, & Bradford, 2013 Purgatory Pyrg Pygulopsis santaclarensis Hershler, Liu, Forsythe, Hovingh &

Wheeler, 2017 Santa Clara Pyrg Pygulopsis santaclaraensis Hershler et al. 2017 JMS 1-11. Recognized Petiton Gastropod 2019-20.

Pyrgulopsis sathos Hershler, 1998 White River Valley Pyrg

Pyrgulopsis saxatilis Hershler, 1998 Sub-globose Snake Pyrg

Pyrgulopsis serrata Hershler, 1998 Northern Steptoe Pyrg

Pygulopsis similis Hershler, Ratcliffe, Liu, Lang, & Hay,

2014 Burnt Corral Pyrg Pyrgulopsis similis Hershler et al. 2014 ZooKeys

429:69-85. Recognized FMCS petition G-2019-21.

Pyrgulopsis simplex Hershler, 1988 Fossil Springsnail

Pyrgulopsis sola Hershler, 1988 Brown Springsnail

Pyrgulopsis stearnsiana (Pilsbry, 1899) Yaqui Springsnail

Pyrgulopsis sterilis Hershler, 1998 Sterile Basin Pyrg

Pyrgulopsis sublata Hershler, 1998 Lake Valley Pyrg

Pyrgulopsis sulcata Hershler, 1998 Southern Steptoe Pyrg

Pyrgulopsis taylori Hershler, 1995 San Luis Obispo Pyrg

Pyrgulopsis texana (Pilsbry, 1935) Phantom Cavesnail

Pyrgulopsis thermalis (Taylor, 1987) New Mexico Hot Springsnail

Pyrgulopsis thompsoni Hershler, 1988 Huachuca Springsnail

Pyrgulopsis torrida Hershler, Liu, Babbitt, Keollogg,

Howard, 2016 Little Sycamore Pyrg

Pyrgulopsis torrida Hershler et al. 2016 Zookeys 601:1-19. Single population likely extinct. Petition Gastropod 2019-22.

Pyrgulopsis transversa Hershler, 1998 Southern Bonneville Pyrg

Pyrgulopsis trivialis (Taylor, 1987) Black River Springsnail

Pyrgulopsis turbatrix Hershler, 1998 Southeast Nevada Pyrg

Pyrgulopsis umbilicata Hershler, 1998 Southern Soldier Meadow Pyrg

Pyrgulopsis variegata Hershler, 1998 Northwest Bonneville Pyrg

Pyrgulopsis varneri Heshler, Liu, & Sada, 2007 Varner's Pyrg

Pyrgulopsis ventricosa Hershler, 1995 Clear Lake Pyrg

Pyrgulopsis villacampae Hershler, 1998 Duckwater Warm Springs Pyrg

Pyrgulopsis vinyardi Hershler, 1998 Vinyards Pyrg

Pyrgulopsis wongi Hershler, 1989 Wong's Pyrg

Rhapinema dacryon Thompson, 1969 Teardrop Snail

Spilochlamys conica Thompson, 1968 Conical Siltsnail

Spilochlamys gravis Thompson, 1968 Armored Siltsnail

(16)

Spilochlamys turgida Thompson, 1969 Pumpkin Siltsnail

Stiobia nana Thompson, 1978 Sculpin Snail

Lithoglyphidae 7 Genera 74 Species

Clappia cahabensis Clench, 1965 Cahaba Pebblesnail

Clappia umbilicata (Walker, 1904) Umbilicate Pebblesnail

Fluminicola ahjumawi Hershler, Liu, Frest & Johannes,

2007 Ahjumawi Pebblesnail

Fluminicola anserinus Hershler, Liu, Frest & Johannes,

2007 Goose Valley Pebblesnail

Fluminicola caballensis Hershler, Liu, Frest & Johannes,

2007 Horse Creek Pebblesnail

Fluminicola coloradoensis Morrison, 1940 Green River Pebblesnail

Fluminicola dalli (Call, 1884) Pyramid Lake Pebblesnail

Fluminicola erosus Hershler, Liu, Frest & Johannes,

2007 Smokey Charley Pebblesnail

Fluminicola favillaceus Hershler, Liu, Frest & Johannes,

2007 Ash Valley Pebblesnail

Flumincola fremonti Hershler, Liu, Frest & Johannes,

2007 Fremont Pebblesnail

Fluminicola fresti Hershler, Liu & Hubbart, 2017 Frest's Pebblesnail

Fluminicola fresti Hershler, Liu, & Hubbart 2017, Zookeys 679:1-20. Recognized Petition Gastropod 2019-2.

Fluminicola fuscus (Haldeman, 1847) Ashy Pebblesnail

Fluminicola gustafsoni Hershler & Liu, 2012 Salmon River Pebblesnail

Fluminicola insolitus Hershler, 1999 Strange Pebblesnail

Fluminicola klamathensis Liu & Hershler, 2019 Klamath Pebblesnail

Fluminicola klamathensis Liu & Hershler 2018 ZooKeys 812:47–67. DOI:

10.3897/zookeys.812.29205. Recognized Petition Gastropod 2019-3.

Fluminicola lunsfordensis Hershler, Liu, Frest & Johannes,

2007 Lunsford Pebblesnail

Fluminicola minutissimus Pilsbry, 1907 Pixie Pebblesnail

Fluminicola modoci Hannibal, 1912 Modoc Pebblesnail

Fluminicola multifarius Hershler, Liu, Frest & Johannes,

2007 Shasta Pebblesnail

Fluminicola neritoides Hershler, Liu, Frest & Johannes,

2007 Willow Creek Pebblesnail

Fluminicola nuttallianus (Lea, 1838) Dusky Pebblesnail

Fluminicola potemicus Hershler, Liu, Frest & Johannes,

2007 Potem Creek Pebblesnail

Fluminicola scopulinus Hershler, Liu, Frest & Johannes,

2007 Castle Creek Pebblesnail

Fluminicola seminalis (Hinds, 1842) Nugget Pebblesnail

Fluminicola turbiniformis (Tryon, 1865) Turban Pebblesnail

(17)

Flumincola umbilicatus Hershler, Liu, Frest & Johannes,

2007 Goose Valley Pebblesnail

Fluminicola umpquaensis Hershler, Liu & Hubbart, 2017 Jade Pebblesnail

Fluminicola umpquaensis Hershler, Liu, Hubbart 2017, Zookeys 679:1-20. Petition Gastropod 2019- 4.

Fluminicola virens (Lea, 1838) Olympia Pebblesnail

Fluminicola virginius Hershler, 1999 Virginia Mountains Pebblesnail Fluminicola warnerensis Hershler, Liu, Frest & Johannes,

2007 Topaz Pebblesnail

Gillia altilis (Lea, 1841) Buffalo Pebblesnail

Lepyrium showalteri (Lea, 1861) Flat Pebblesnail

Pristinicola hemphilli (Pilsbry, 1890) Pristine Pyrg

Somatogyrus alcoviensis Krieger, 1972 Reverse Pebblesnail

Somatogyrus amnicoloides Walker, 1915 Ouachita Pebblesnail

Somatogyrus aureus Tryon, 1865 Golden Pebblesnail

Somatogyrus biangulatus Walker, 1906 Angular Pebblesnail

Somatogyrus constrictus Walker, 1904 Knotty Pebblesnail

Somatogyrus coosaensis Walker, 1904 Coosa Pebblesnail

Somatogyrus crassilabris Walker, 1915 Thick-lip Pebblesnail

Somatogyrus crassus Walker, 1904 Stocky Pebblesnail

Somatogyrus currierianus (Lea, 1863) Tennessee Pebblesnail

Somatogyrus decipiens Walker, 1909 Hidden Pebblesnail

Somatogyrus depressus (Tryon, 1862) Sandbar Pebblesnail

Somatogyrus excavatus Walker, 1906 Ovate Pebblesnail

Somatogyrus georgianus Walker, 1904 Cherokee Pebblesnail

Somatogyrus hendersoni Walker, 1909 Fluted Pebblesnail

Somatogyrus hinkleyi Walker, 1904 Granite Pebblesnail

Somatogyrus humerosus Walker, 1906 Atlas Pebblesnail

Somatogyrus integra (Say, 1829) Ohio Pebblesnail

Somatogyrus nanus Walker, 1904 Dwarf Pebblesnail

Somatogyrus obtusus Walker, 1904 Moon Pebblesnail

Somatogyrus parvulus Tryon, 1865 Sparrow Pebblesnail

Somatogyrus pennsylvanicus Walker, 1904 Shale Pebblesnail

Somatogyrus pilsbryanus Walker, 1904 Tallapoosa Pebblesnail

Somatogyrus pumilus (Conrad, 1834) Compact Pebblesnail

Somatogyrus pygmaeus Walker, 1909 Pygmy Pebblesnail

(18)

Somatogyrus quadratus Walker, 1906 Quadrate Pebblesnail

Somatogyrus rheophilus Thompson, 1984 Flint Pebblesnail

Somatogyrus rosewateri Gordon, 1986 Elk Pebblesnail

Somatogyrus sargenti Pilsbry, 1895 Mud Pebblesnail

Somatogyrus strengi Pilsbry & Walker, 1906 Rolling Pebblesnail

Somatogyrus substriatus Walker, 1906 Choctaw Pebblesnail

Somatogyrus tenax Thompson, 1969 Savannah Pebblesnail

Somatogyrus tennesseensis Walker, 1906 Opaque Pebblesnail

Somatogyrus trothis Doherty, 1878

Somatogyrus tryoni Pilsbry & Baker, 1927 Coldwater Pebblesnail

Somatogyrus virginicus Walker, 1904 Panhandle Pebblesnail

Somatogyrus walkerianus Aldrich, 1905 Gulf Coast Pebblesnail

Somatogyrus wheeleri Walker, 1915 Channeled Pebblesnail

Taylorconcha insperata Hershler, Liu, Frest, Johannes, &

Clark, 2006 Unexpected Pebblesnail

Taylorconcha serpenticola Hershler, Frest, Johannes, Bowler, &

Thompson, 1994 Bliss Rapids Snail

Family Pleuroceridae 7 Genera 165 Species

Athearnia anthonyi (Redfield, 1854) Anthony's Riversnail

Athearnia crassa (Haldeman, 1842) Boulder Snail

Elimia acuta (I. Lea, 1831) Acute Elimia

Elimia alabamensis (I. Lea, 1861) Mud Elimia

Elimia albanyensis (I. Lea, 1864) Black-crest Elimia

Elimia ampla (J. G. Anthony, 1854) Ample Elimia

Elimia annae Mihalcik & F. G. Thompson, 2002 Rainbow Elimia

Elimia annettae (Goodrich, 1941) Lilyshoals Elimia

Elimia arachnoidea (Anthony, 1854) Spider Elimia

Elimia aterina (I. Lea, 1863) Coal Elimia

Elimia athearni (Clench & Turner, 1956) Knobby Elimia

Elimia bellacrenata (Haldeman, 1842) Princess Elimia

Elimia bellula (I. Lea, 1861) Walnut Elimia

Elimia boykiniana (Lea, 1840) Flaxen Elimia

Elimia brevis (I. Lea, 1843) Short Spire Elimia

Elimia broccata Thompson, 2000 Brooch Elimia

Elimia buffyae Mihalcik & F. G. Thompson, 2002 Iris Elimia

(19)

Elimia bullula (I. Lea, 1861) Yellowleaf Elimia

Elimia caelatura (Reeve, 1860) Savannah Elimia

Elimia cahawbensis (I. Lea, 1861) Cahaba Elimia

Elimia capillaris (I. Lea, 1861) Spindle Elimia

Elimia carinifera (Lamarck, 1822) Sharp-crest Elimia

Elimia carinocostata (Lea, 1854) Fluted Elimia

Elimia catenaria (Say, 1822) Gravel Elimia

Elimia catenoides (I. Lea, 1842) Lirate Elimia

Elimia chiltonensis (Goodrich, 1941) Prune Elimia

Elimia christyi (I. Lea, 1862) Christy's Elimia

Elimia clara (Anthony, 1854) Riffle Elimia

Elimia clausa (I. Lea, 1861) Closed Elimia

Elimia clavaeformis (I. Lea, 1841) Club Elimia

Elimia clenchi (Goodrich, 1924) Slackwater Elimia

Elimia cochliaris (I. Lea, 1868) Cockle Elimia

Elimia comalensis (Pilsbry, 1890) Balcones Elimia

Elimia comma (Conrad, 1834) Hispid Elimia

Elimia costifera (Reeve, 1861) Corded Elimia

Elimia crenatella (I. Lea, 1860) Lacy Elimia

Elimia curreyana (I. Lea, 1841) Amber Elimia

Elimia cylindracea (Conrad, 1834) Cylinder Elimia

Elimia darwini Mihalcik & F. G. Thompson, 2002 Pup Elimia

Elimia dickinsoni (Clench & Turner, 1956) Stately Elimia

Elimia dislocata (Reeve, 1861) Lapped Elimia

Elimia dooleyensis (I. Lea, 1862) Graphite Elimia

Elimia ebenum (I. Lea, 1841) Ebony Elimia

Elimia edgariana (I. Lea, 1841) Cumberland Elimia

Elimia exusta Mihalcik & F. G. Thompson, 2002 Fire Elimia

Elimia fascinans (I. Lea, 1861) Banded Elimia

Elimia flava (I. Lea, 1862) Yellow Elimia

Elimia floridensis (Reeve, 1860) Rasp Elimia

Elimia fusiformis (I. Lea, 1861) Fusiform Elimia

Elimia gibbera (Goodrich, 1922) Shouldered Elimia

Elimia glarea Mihalcik & F. G. Thompson, 2002 Gravel Elimia

Elimia godwini F. G. Thompson, 2000 Rusty Elimia

(20)

Elimia hartmaniana (I. Lea, 1861) High-spired Elimia

Elimia haysiana (I. Lea, 1843) Silt Elimia

Elimia hydei (Conrad, 1834) Gladiator Elimia

Elimia impressa (I. Lea, 1841) Constricted Elimia

Elimia inclinans (I. Lea, 1862) Slanted Elimia

Elimia interrupta (Haldeman, 1840) Knotty Elimia Accidental deletion

Elimia interveniens (I. Lea, 1862) Slowwater Elimia

Elimia induta (I. Lea, 1862) Gem Elimia

Elimia jonesi (Goodrich, 1936) Hearty Elimia

Elimia lachryma (Reeve, 1861) Teardrop Elimia

Elimia laeta (Jay, 1839) Ribbed Elimia

Elimia laqueata (Say, 1829) Panel Elimia

Elimia lecontiana (I. Lea, 1841) Rippled Elimia

Elimia livescens (Menke, 1830) Liver Elimia

Elimia macglameriana (Goodrich, 1936) Wrinkled Elimia

Elimia melanoides (Conrad, 1834) Black Mudalia

Elimia mihalcikae F. G. Thompson, 2000 Latticed Elimia

Elimia modesta (I. Lea, 1845) Coldwater Elimia

Elimia mutabilis (I. Lea, 1862) Oak Elimia

Elimia nassula (Conrad, 1834) Round-ribbed Elimia

Elimia olivula (Conrad, 1834) Caper Elimia

Elimia ornata (I. Lea, 1868) Ornate Elimia

Elimia paupercula (I. Lea, 1862) Sooty Elimia

Elimia perstriata (I. Lea, 1852) Engraved Elimia

Elimia pilsbryi (Goodrich, 1927) Rough-lined Elimia

Elimia plicatastriata (Wetherby, 1876) Carved Elimia

Elimia porrecta (I. Lea, 1863) Nymph Elimia

Elimia potosiensis (I. Lea, 1841) Pyramid Elimia

Elimia proxima (Say, 1825) Sprite Elimia

Elimia pupaeformis (I. Lea, 1864) Pupa Elimia

Elimia pupoidea (J. G. Anthony, 1854) Bot Elimia

Elimia pybasii (I. Lea, 1862) Spring Elimia

Elimia pygmaea (Smith, 1936) Pygmy Elimia

Elimia semicarinata (Say, 1829) Fine-ridged Elimia

(21)

Elimia shenandoa (Dillon, 2019) Blue Ridge Elimia

Pleurocera shenandoa Dillon, 2019 was recoginzed as Elimia shenandoa (Dillon, 2019). FMCS petition G-2021-20

Elimia showalterii (I. Lea, 1860) Compact Elimia

Elimia simplex (Say, 1825) Smooth Elimia

Elimia striatula (I. Lea, 1842) File Elimia

Elimia strigosa (I. Lea, 1841) Brook Elimia

Elimia symmetrica (Haldeman, 1841) Symmetrical Elimia

Elimia taitiana (I. Lea, 1841) Dented Elimia

Elimia teres (I. Lea, 1841) Elegant Elimia

Elimia teretria F. G. Thompson, 2000 Auger Elimia

Elimia timida (Goodrich, 1942) Timid Elimia

Elimia troostiana (I. Lea, 1838) Mossy Elimia

Elimia ucheensis (I. Lea, 1862) Creek Elimia

Elimia vanhyningiana (Goodrich, 1921) Goblin Elimia

Elimia vanuxemiana (I. Lea, 1843) Cobble Elimia

Elimia varians (I. Lea, 1861) Puzzle Elimia

Elimia variata (I. Lea, 1861) Squat Elimia

Elimia viennaensis (I. Lea, 1862) Slough Elimia

Elimia virginica (Gmelin, 1791) Piedmont Elimia

Gyrotoma excisa (I. Lea, 1843) Excised Slitshell

Gyrotoma lewisii (I. Lea, 1869) Striate Slitshell

Gyrotoma pagoda (I. Lea, 1845) Pagoda Slitshell

Gyrotoma pumila (I. Lea, 1860) Ribbed Slitshell

Gyrotoma pyramidata Shuttleworth, 1845 Pyramid Slitshell

Gyrotoma walkeri (H. H. Smith in Goodrich, 1924) Round Slitshell

Io fluvialis (Say, 1825) Spiny Riversnail

Leptoxis ampla (J. G. Anthony, 1855) Round Rocksnail

Leptoxis arkansensis (Hinkley, 1915) Arkansas Mudalia

Leptoxis carinata (Bruguière, 1792) Crested Mudalia

Leptoxis clipeata (H. H. Smith in Goodrich, 1922) Agate Rocksnail

Leptoxis compacta (J. G. Anthony, 1854) Oblong Rocksnail

Leptoxis coosaensis (I. Lea, 1861) Painted Rocksnail

Whelan et al., 2017 Zookeys 697:21-36. Revised taxonomy of Leptoxis taeniata. Petition Gastropod 2019-07.

Leptoxis dilatata (Conrad, 1835) Seep Mudalia

(22)

Leptoxis foremani (I. Lea, 1843) Interrupted Rocksnail

Leptoxis formosa (I. Lea, 1860) Maiden Rocksnail

Leptoxis ligata (J. G. Anthony, 1860) Rotund Rocksnail

Leptoxis lirata (H. H. Smith in Goodrich, 1922) Lirate Rocksnail

Leptoxis minor (Hinkley, 1912) Knob Mudalia

Leptoxis occultata (H. H. Smith in Goodrich, 1922) Bigmouth Rocksnail

Leptoxis picta (Conrad, 1834) Spotted Rocksnail

Leptoxis plicata (Conrad, 1834) Plicate Rocksnail

Leptoxis praerosa (Say, 1821) Onyx Rocksnail

Leptoxis showalterii (I. Lea, 1860) Coosa Rocksnail

Leptoxis torrefacta (H. H. Smith in Goodrich, 1922) Squat Rocksnail

Leptoxis trilineata (Say, 1829) Broad Mudalia

Leptoxis umbilicata (Wetherby, 1876) Umbilicate Rocksnail

Leptoxis virgata (I. Lea, 1841) Smooth Mudalia

Leptoxis vittata (I. Lea, 1860) Striped Rocksnail

Lithasia armigera (Say, 1821) Armored Rocksnail

Lithasia bubala Minton, 2013 Buffalo Rocksnail Lithasia bubala Minton, 2013, The Nautilus 127(3)

:119-124. Recognized FMCS petition G-2019-8.

Lithasia curta (I. Lea, 1868) Knobby Rocksnail

Lithasia duttoniana (I. Lea, 1841) Helmet Rocksnail

Lithasia geniculata (Haldeman, 1840) Ornate Rocksnail

Lithasia hubrichti Clench, 1956 Big Black Rocksnail

Lithasia jayana (I. Lea, 1841) Rugose Rocksnail

Lithasia lima (Conrad, 1834) Warty Rocksnail

Lithasia obovata (Say, 1829) Shawnee Rocksnail

Lithasia salebrosa (Conrad, 1834) Muddy Rocksnail

Lithasia spicula Minton, Savarese, & D. C. Campbell,

2005 Harpeth Rocksnail

Lithasia verrucosa (Rafinesque, 1820) Varicose Rocksnail

Pleurocera acuta Rafinesque in Blaineville, 1824 Sharp Hornsnail

Pleurocera alveare (Conrad, 1834) Rugged Hornsnail

Pleurocera annulifera (Conrad, 1834) Ringed Hornsnail

Pleurocera brumbyi (I. Lea, 1852) Spiral Hornsnail

Pleurocera canaliculata (Say, 1821) Silty Hornsnail

Pleurocera corpulenta (J.G. Anthony, 1854) Corpulent Hornsnail

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Pleurocera curta (Haldeman, 1841) Shortspire Hornsnail

Pleurocera foremani (I. Lea, 1843) Rough Hornsnail

Pleurocera gradata (J. G. Anthony, 1854) Bottle Hornsnail

Pleurocera nobilis (I. Lea, 1845) Noble Hornsnail

Pleurocera parva (I. Lea, 1862) Dainty Hornsnail

Pleurocera postelli (I. Lea, 1862) Broken Hornsnail

Pleurocera prasinata (Conrad, 1834) Smooth Hornsnail

Pleurocera pyrenella (Conrad, 1834) Skirted Hornsnail

Pleurocera showalteri (I. Lea, 1862) Upland Hornsnail

Pleurocera striatum (I. Lea, 1863) Striate Hornsnail

Pleurocera trochiformis (Conrad, 1834) Sulcate Hornsnail

Pleurocera uncialis (Reeve, 1861) Pagoda Hornsnail

Pleurocera vestita (Conrad, 1834) Brook Hornsnail

Pleurocera walkeri Goodrich, 1928 Telescope Hornsnail

Family

Semisulcospiridae 1 Genus 11 Species

Juga acutifilosa (Stearns, 1890) Topaz Juga

Juga bulbosa (Gould, 1847) Bulb Juga

Juga chacei (Henderson, 1935) Chace Juga

Juga hemphilli (Henderson, 1935) Barren Juga

Juga interioris (Goodrich, 1944) Smooth Juga

Juga laurae (Goodrich, 1944) Oasis Juga

Juga newberryi (Lea, 1860) Banded Juga

Juga nigrina (Lea, 1856) Black Juga

Juga occata (Hinds, 1844) Scalloped Juga

Juga plicifera (Lea, 1838) Pleated Juga

Juga silicula (Gould, 1847) Glassy Juga

Family Pomatiopsidae 1 Genus 6 Species

Pomatiopsis binneyi Tryon, 1863 Robust Walker

Pomatiopsis californica Pilsbry, 1899 Pacific Walker

Pomatiopsis chacei Pilsbry, 1937 Marsh Walker

Pomatiopsis cincinnatiensis (I. Lea, 1840) Brown Walker

Pomatiopsis hinkleyi Pilsbry, 1896 Tennessee River Walker

Pomatiopsis lapidaria (Say, 1817) Slender Walker

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Family Valvatidae 1 Genus 10 Species

Valvata bicarinata Lea, 1841 Two-ridge Valvata

Valvata humeralis Say, 1829 Glossy Valvata

Valvata lewisi Currier, 1868 Fringed Valvata

Valvata mergella Westerlund, 1883 Rams-horn Valvata

Valvata perdepressa Walker, 1906 Purplecap Valvata

Valvata sincera Say, 1824 Mossy Valvata

Valvata tricarinata (Say, 1817) Threeridge Valvata

Valvata utahensis Call, 1884 Desert Valvata

Valvata virens Tryon, 1863 Emerald Valvata

Valvata winnebagoensis Baker, 1928 Flanged Valvata

Taxa included in Johnson et al., 2013 but no longer recognized due to synonymy.

Species Author AFS Common Name Change Notes

Lanx sp. Clark, Campbell & Lydeard, 2017 Bliss Rapids Snail Formal description completed for the Bliss Rapids Snail. Petition: Gastrpoda-2019-05

Lymnaea atkaensis (Dall, 1884) Frigid Lymnaea Placed in Ladislavella. Petition Gastropod 2021-02

Lymnaea producta (Mighels, 1845) Uncertain Classification Placed in Pseudisidora. Petition Gastropod 2021-03

Lymnaea rubella (I. Lea, 1841) Aloha Lymnaea Placed in Pseudisidora. Petition Gastropod 2021-03

Stagnicola apicina (I. Lea, 1838) Abbreviate Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola arctica (I. Lea, 1864) Artic Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola bonnevillensis (Call, 1884) Fat-Whorled Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola caperata (Say, 1829) Wrinkled Marshsnail Placed in Hinkleyia. Petitition Gastropod 2021-01 Stagnicola catascopiium (Say, 1817) Woodland Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola contracta (Currier in DeCamp, 1881) Deepwater Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02

Stagnicola elodes (Say, 1821) Marsh Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02

Stagnicola elrodi (F.C. Baker & Henderson, 1933) Flathead Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola emarginata (Say, 1821) St. Lawrence Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02

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Stagnicola exilis (I. Lea, 1834) Flat-Whorled Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola gabbii (Tryon, 1865) Striate Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola hinkleyi (F.C. Baker, 1906) Rustic Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola idahoensis (Henderson, 1931) Shortspire Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola kennicotti (F.C. Baker, 1933) Western Artic Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola mighelsi (Binney, 1865) Bigmouth Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola neopalustris (F.C. Baker, 1911) Piedmont Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola oronoensis (F.C. Baker, 1904) Obese Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola petoskeyensis (B. Walker, 1908) Petoskey Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola pilsbryi (Hemphill, 1890) Fish Springs Marshsnail Placed in Hinkleyia. Petitition Gastropod 2021-01 Stagnicola traskii (Tryon, 1865) Widelip Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola utahensis (Call, 1884) Thickshell Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02 Stagnicola woodruffi (F.C. Baker, 1913) Coldwater Pondsnail Placed in Ladislavella. Petition Gastropod 2021-02

Archiphysa ashmundi Taylor, 2003 San Rafel Physa Archiphysa placed in Physella. Petition Gastropod-

2019-01 and adopted 2021

Archiphysa sonomae Taylor, 2003 Sonoma Physa

Archiphysa placed in Physella. Petition Gastropod- 2019-01 and adopted 2021

Laurentiphysa chippuvarum Taylor, 2003 Chippewa Physa

Laurentiphysa placed in Physa. Petition Gastropod 2019-02

Physella mexicata (Philippi in Küster, 1841) Polished Physa

Synoymized with P. acuta (Drparnaud, 1805).

Petition Gastropod 2021-08

Physella virgata (Gould, 1855) Protean Physa

Synonymized with P. acuta (Drparnaud, 1805).

Petition Gastropod 2021-08

Physella winnipegensis Pip, 2004 Lake Winnipeg Physa

Synonymized with P. acuta (Draparnaid, 1805).

Petition Gastropod 2021-07

Ferrissia fragilis (Rowell, 1863) Fragile Ancylid

Ferrissia californica (Rowell, 1863) is the priority name for F. fragilis. Petition Gastropod 2021-09

Micromenetus brogniartianus (Lea, 1842) Disc Sprite

Micomenetus placed in Dilatata. Petition Gastropod 2019-09. Petition tabled but adopted for Dilatata 2021.

Micromenetus dilatatus (Gould, 1841) Bugle Sprite

Micomenetus placed in Dilatata. Petition Gastropod 2019-09. Petition tabled but adopted for Dilatata 2021.

Micromenetus floridensis (Baker, 1945) Penny Sprite

Micomenetus placed in Dilatata. Petition Gastropod 2019-09. Petition tabled but adopted for Dilatata 2021.

Micromenetus sampsoni (Ancey, 1885) Sampson Sprite

Micomenetus placed in Dilatata. Petition Gastropod 2019-09. Petition tabled but adopted for Dilatata 2021.

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Viviparus georgianus (Lea, 1834) Banded Mysterysnail

New combination Stelbrink et al. 2020. Petition Gastropod 2021-13.

Viviparus goodrichi (Archer, 1933) Globose Mysterysnail

New combination Stelbrink et al. 2020. Petition Gastropod 2021-13.

Viviparus intertextus (Say, 1829) Rotund Mysterysnail

New combination Stelbrink et al. 2020. Petition Gastropod 2021-13.

Viviparus limi Pilsbry, 1918 Ochlockonee Mysterysnail

New combination Stelbrink et al. 2020. Petition Gastropod 2021-13.

Vivparus subpurpurus (Say, 1829) Olive Mysterysnail

New combination Stelbrink et al. 2020. Petition Gastropod 2021-13.

Assiminea infima Berry, 1941 Badwater Snail

Placed in Angustassiminea Habe, 1943. Petition Gastropod 2021-14

Leptoxis taeniata (Conrad, 1834) Painted Rocksnail Leptoxis coosaensis first available name. Petition Gastropoda 2019-07

Figure

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References

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