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Prediction of Life History Traits in Invertebrate Species Exposed to Pesticide

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bHbHb bMbHb bHbHb

Ž 6 Ž bMbHb bHbHb bHbHb bHbHb # # # # # # bHbHb Ž Ž Ž 6 c " Ž eTf f f f f f f f f f f f f f g

(17)

i,40)X ')(+N-')(+?)(I5d0),/5-l 02X-(m8;(I.+b-5-*-,}0FA6 6 '@(IN-'@(I?@(I5d02,45-l 0)X-(cN-')39J[D9J-,4g/,40FA 023 ?@b-'@GO,4G9(DL5-*

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?)bK'@GO,4G936')?@X-,/N=8;39')e†b-g/D9?8;36'"02X-(:Nk(qDEDLN-X-,/*T(1QNk(I'@,/eT(I5d02?j.qD95xJP(:8;36b-5-*x,/5TRD9JKg/( 6#9#hX-(+A

D9'@(EJ[D9?@(I*m365mNk(I'@.I(+5H0ex36'0fD9g4,40FA€DL5-*ƒNk(I').+(I5d0g/39?)?‚'@(IN-'@3*Kb-.+0),/365SG9DLg/b-(+?‚iX-,/.fXio(+')(*-(…{

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|8io(\_H5-3ui 02XK(D9l6(>*-,/?02'@,/J-bK0),/365x36'rNP36NKb-gDL0),/365T*-(+5-?),}0FA DL0D9l6( MDL5-*=02,4ex( …ij(.ID95

N-'@3y@(I.10j0)3Z0),/eT( 6#&hXK(I')(18;36')(LNk36N-bKgDL0),/365nl6'@3 ih0)Xx.ID95=Jk(h8;36b-5-*TGO,D\')(+NP(IDL02(+*=exDL02'@,}Q

e†b-g}02,4N-g/,4.qDL0),/365-?CD9?"?)XK3 i5x,/5T &wHb[DL02,4365 6#ŒO#R|0r.qD95xD9g4?)3JP(i'),}0)0)(I5TD9?(

8

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_` ` ` ` ` ` ` ` ` ` a ( 8 ( 8 # # # c "; ( 8 eTf f f f f f f f f f g _` ` ` ` ` ` ` ` ` ` ` ` ` ` a bMbHb dc "; 6 Ž

bHbHb bMbHb bHbHb

Ž 6 Ž bMbHb bHbHb bMbHb bHbHb # # # # # # bHbHb

Ž Ž Ž 6 c

" Ž ef f f f f f f f f f f f f f g _` ` ` ` ` ` ` ` ` ` a ( ( # # # c "; ( eTf f f f f f f f f f g b

U 6#Œ9`

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02,}G6(S'2DL0)(U =

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Nk36N-b-g/DL02,4365{Fg4(+G9(Ig‹(+.I390)3qQK,4.I36g436l6,4.qD9gk(I5-*KNP36,45d02?M",45H0)'),45-?),4.\'2DL0)(\398R,45-.I'@(qD9?@(xU `D95-*S*-,/'@(I.10

,/5K?@0fDL5H02D95-(+36b-?l6'@3uih02XY'2D02(U ML;`…#"]o390)X ex(ID9?)bK')(IeT(I5d0)?hD9')(>398„,/5d0)(I')(+?@00)3†(I.+36g/39l6,/?02?I#RhX-(

(18)

D9l6(\*K,/?@0)'),4J-bK02,4365sKD9g/gs3L8CiX-,4.fXNK')3uGO,/*-(eT36')(\,45-?),4l6Xd0h,/5d0230)X-(NP36NKb-gDL0),/365=*KAO5[D9eT,/.I?+#

32 5 4 9 4 0(9 59

[36'‚39b-'\b-5-*-(+')?0fD95-*K,/5-l-Pio(nb-?@(I* &] 023Y'@(I.+36exNKbK02(nD9g4gC3L8r0)X-(†(+.I36g436l6,/?02?+~s(I?@g/,/(

?),4e†b-g/DL02,4365-?I#‰B($i,4g/g*-,4?).+b-?)?39b-'‚?@,/e†b-g/DL02,4365-?:D95-*')(+?)b-g}02?I#‚ RQKNk(I'@,/eT(I5d0fDLgC*[D0fDT8;39'>02XK(

.I395H0)')36g(…QNP(+'),/eT(I5d0i,40)XSD9N-X-,4*-?h(1QNk36?)(+*mD9?h5-(+365[DL0)(I?:,/?hl6,}G6(I5S,/5€CD9J-g4( 6#ŒO#rhXK,/?h*[DL02D

ioD9?nb-?)(+*shD9g4365-lƒi,40)X D95 D9?)?@b-ex(+*Nk(I'@.I(I5d0xex39'@0fDLg/,40FABD95K* Nk(I'@.I(+5H0xg/39?)?')(+N-')3O*-b-.102,4365

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CD9J-g/( 6#aŒJro365d0)')36g„\DL02DE8;36'!>(+365[DL0)(Z QNk36?)bK')(

+- > ' $ = C

EAB ;I :dŽ :dŽ )< 6Œ

EAB EAO L Ž Ž Ž Ž Ž Ž Œ ':HŒ ŒLŽ Œ *<

+- > ' $ $$ $= $ $ $ $ $DC $ $

EAB ;I 9Ž 6Ž 6Ž Œ Œ Œ3<

EAB EAO L "<9Ž Œ9ŽdŒ Œ9Ž ŒLŽd ':(: dŒ qŽ

+->

'

=

=B$ == = = =

=

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EAB ;I Œ(< Œ Œ6Œ Œ u Œ uŽ  

EAB EAO L < : )< Œ2: Œ9Ž 

+->

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B$ =

1C

EAB ;I   Œ Œ

EAB EAO L Œ Œ Ž Ž Ž Ž Ž Ž

,/eT,/g/D9'†(…QKNk(I'@,/eT(I5d0fDLgh*[DL0fDc8;36'TD9g/gh?@,}Q .+365-.I(+5d02'2D02,/395-?x3L8\D9N-XK,/*-?T(1QNk36?)(+* JP3L02X DL?

(19)

8;(I.+b-5-*-,}0FA6O8;36'j(ID9.fX .+365-.I(+5d02'2D02,/395=.qDL5 JP(8;36b-5K* ,/5TRDLJ-g/( 6#ED95-*x,/5 R,/l6bK')( 6#6#R‰B(‚?)(+(

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†K02X-D95Sij(I'@(Z0)X-(ZD9*-b-g}02?+#

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uŽ9N-NKe < "<  Œ

Œ9Ž9N-NKe 9Ž Œ : d :dŽ

:dŽ9N-NKe <LŽ 6Œ )< 9Ž

6Ž9N-NKe 9Ž 6  :

6Ž9N-NKe 6 6 uŽ9Ž d

uŽ6Ž9N-NKe dŒ <9Ž

0

10

20

30

40

50

60

70

80

0

10

20

30

40

50

60

70

80

90

100

Concentration−Mortality Curve

Concentration Exposure

Percent Mortality

Neonate

Adult

0

10

20

30

40

50

60

70

80

0

10

20

30

40

50

60

70

80

90

100

Concentration−Loss Reproduction Curve

Concentration Exposure

Percent Loss Reproduction

Neonate

Adult

R,4l6b-'@( 6# "%C(I'@.I(+5H0 36'0fD9g4,40FAmUg/(18ˆ0…`oDL5-*M%p(I'@.I(I5d0„39?)?::(+N-')3O*-b-.102,4365BU'),4l6Xd0…`

(20)

5-(+365[DL0)(I?hiDL?:'@(I*-bK.I(I*MD9?h.I395-.I(+5H0)'2DL0),/365€398 MDL')l636?)D95{ ,/5K.I')(ID9?)(+*s# [36'hD95€(1QNk36?)bK')(398

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Œ :c*[DqAO?n.I36eTN[D9'@(I* i,402X dŒƒ*[DqAO?8;')36e 02X-(€.+365d02')39g #7hX-(+?)(')(I?@b-g40)?nD9')(€?@b-N-Nk36'@0)(I* JHA

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MD9'@l636?)D95{ ,/5 D‡.I365K.I(I5d0)'2DL0),/365{|*-(INk(I5K*-(I5d0 exD95-5K(I'I# >g/?@3-o,}0x?@X-36b-g4*Jk(M5-3L02(I* 02X[D0

b-5-*K(I'†*-, ‹(+')(+5H0E.I365K.I(I5d0)'2DL0),/365-?+R02XK( ( P(I.10†398h0)X-(=,45-?)(+.+02,4.I,4*-(=*-(+.Ig4,/5-(+?†D9?$36g/*-(+'EDLN-X-,/*K?

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0

5

10

15

20

25

30

35

0

0.1

0.2

0.3

0.4

0.5

0.6

0.7

0.8

0.9

1

Survival of A. Pisum Neonates Exposed to Margosan−O

Percent Survival

Aphid Age

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

0

5

10

15

20

25

30

35

0

0.1

0.2

0.3

0.4

0.5

0.6

0.7

0.8

0.9

1

Survival of A. Pisum Adults Exposed to Margosan−O

Percent Survival

Aphid Age

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

100 ppm

R,4l6b-')( 9#ŒJrOb-'@GO,4GLD9g‹398!:(I365-DL02(Z RQNP36?@b-')(=Ug4(+8ˆ0f`D95-*S‚*-b-g}0: RQKNk36?@b-')(=U'@,/l6Xd0…`

hX-( '@(IgD02,/395-?)X-,4NBJk(+0Fij(I(+5 .I365-.+(I5d02')DL02,4365D95K*8;(I.+b-5-*-,}0FA,4?E?),4ex,4g D9'>023M0)X[DL0†3L8h.I365O{

(21)

MD9'@l636?)D95{ D9?n5-(I395[DL02(+?nb-5-*-(+')l63BD‡XHb-l9(S*K(I.I'@(qD9?@(M,45702X-(+,/'†8;(I.+b-5-*-,}0FA '2DL0)(I?+# [36'nDL?

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D95S(1QNk36?)b-'@(E398 uŽ6Ž9NKN-e.I36eTN[D9'@(I*€i,40)XMDne=D QK,4e†b-e 8;(I.IbK5-*-,40FAS398D9')39b-5-* 6Ž 8;36'h02XK(

.I395H0)')36gW#

0

5

10

15

20

25

30

35

0

1

2

3

4

5

6

7

8

Fecundity of A. Pisum Neonates Exposed to Margosan−O

Fecundity

Aphid Age

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

0

5

10

15

20

25

30

35

0

1

2

3

4

5

6

7

8

9

Fecundity of A. Pisum Adults Exposed to Margosan−O

Fecundity

Aphid Age

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

100 ppm

R,4l6b-'@( 6#J ^(+.Ib-5K*-,40FAY3L8!:(I365[D02(Z RQKNk36?@b-')(=Ug4(+8ˆ0…`DL5-*>*-b-g}0: QNk36?)bK')(=U'),4l6Xd0…`

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(22)

(I?@g/,/(exDL02'@,}Q=023T36JK0fDL,/5€36b-'Nk36N-b-g/DL02,4365YN-')3 y@(I.+0),/365K?I#>g/gs?@,/e†b-g/DL02,4365-?oij(I'@(E'@b-58;36' uŽ9Ž

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0

10

20

30

40

50

60

70

80

90

100

0

1

2

3

4

5

6

7

8

x 10

18

Popluation Structure

Time

Number of Individuals

(23)

'2D02(‡U4=

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(1QNk36?)bK')(9# ')D9N-X-?3L8 =

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D=l6(+5-(I')DL02,4365s#‚‰B(†?@(I(†02X-DL0 =

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b

"

8;36' 6Ž9N-N-e (1QNk36?)bK')(9# [36'

D9*-bKg40"(1QNk36?)bK')(9O02X-(5-(10o'@(IN-'@3*Kb-.+0),4G6(:'2D02(hiDL? 9#8;36'&02XK(:.+365d02'@36gkD95-*=Œ#<‚8;36' uŽ9Ž9N-N-e

(1QNk36?)bK')(9#EhX-(')(+*-b-.+0),/365c398^=

ioD9?5-390DL?\?)b-J-?0fD95d0),D9gRi,}02XcD9*Kb-g40\(…QNP36?@b-')(L„J-bK0\02XK(

( P(I.10‚3L8C0)X-(.fX-(IeT,/.ID9gs.qDL5?02,/g4gsJk(Z?@(I(+5s# S(qD95Sl6(I5K(I'2D02,/395€0),/eT(TU@‚`D9g4?)3n*-(I.+g/,45-(I*Yi,40)X

,/5K.I')(ID9?)(+*NP(+?@0),/.I,4*-(=.+365-.+(I5d02')DL02,4365s#MhXK(

ex39')(=Nk(I?02,4.I,/*K(T0)X-(YD9N-XK,/*-?Zio(+')(Y(1QNk36?)(+*7023-02XK(xg43 ij(I'\0)X-(nex(ID95Bl9(I5-(+'2DL0),/365m02,4ex(L#T<>3uio(1G6(+'IR0)X-(=DLN-X-,/*K?Z(…QNP36?@(I*‡8;')36e JK,/'@0)X‡ij(I'@(

D# P(I.102(I*SeT36')(0)X[D95€02X-(ZD9N-XK,/*-?h(…QKNk36?@(I*ƒDL?>DL*-b-g40)?8;36'D9g4gsNP(+?@0),/.I,4*-(\.I395-.I(+5H0)'2DL0),/365K?I#

0

10

20

30

40

50

60

70

80

0

10

20

30

40

50

60

70

80

90

Net Reproductive Rate

Concentration Exposure

Net Reproductive Rate (R

0

)

Neonate

Adult

0

10

20

30

40

50

60

70

80

8

9

10

11

12

13

14

Mean Generation Time

Concentration Exposure

Generation Time (T)

Neonate

Adult

R,4l6b-')( 6#aJ!>(10 (IN-'@3*Kb-.+0),4G6(:DL02(SD95-* (qDL5 \(I5-(+'2DL0),/365 h,/eT( 8;36'T!>(+365[DL0)(I? DL5-*

(24)

5-.I(‚ij(36J0fD9,45-(I*=0)X-(>GLD9g4b-(I?j398 =

DL5-* @Zij(‚ij(I')(D9J-g/(023E*-(+0)(I')eT,/5K(>02XK(‚,/5d0)'),/5K?),/.

'2D02(=3L8,/5-.+')(qDL?)(cU o`Z8;36'Z(qD9.fX (1QNk36?)bK')( .ID9?)(L#m‚.I.+36')*K,/5-lM0)3 ‰ p02X-(=,/5d02'@,/5-?@,/.T'2DL0)(

398:,/5K.I')(ID9?)(Y,4?DƒeT(qD9?@b-')(Y3980)X-(€D9J-,4g/,40FAc3L8‚DƒNk36N-bKg DL0),/365‡023m,/5K.I')(ID9?)(Y(…QNP365K(I5d02,/D9g/g}A,45

D95 b-5-g4,/eT,40)(I* (+5HGO,4')365-eT(I5d0I# hXHb-? ,4?€D95 ,/eTNP36'0fD95d0=ex(ID9?)b-'@(IeT(I5d0S398$NP36NKb-g DL0),/365

*KAO5[D9eT,/.+?I#xhX-,/?Z'2DL0)(n,/?l6,4G9(I5‡JdA

GJI

K

# b-'.qD9g4.Ib-g/DL02(+* GLD9g4b-(I?ZD9'@(n*-(IN-,4.+0)(I*

,/5 C,/l6b-'@( 6#K# [36' 5-(+365[DL0)(m(1QNP39?)b-'@(9 *-(I.+')(ID9?)(+* ,45 D7.I365-.+(I5d02')DL02,4365 *-(INk(I5K*-(I5d0

exD95-5-(+'I^JP(+.I36eT,/5KlT5K(IldDL0),4G9(EDL0 :dŽ9NKN-e#jv5K(IldDL0),4G9( ,4exNKg/,/(+?hNP39N-b-gD02,/395Y(1QO02,45-.+0),/365s#

F5d02(+')(I?02,45-l6g4A98;36'D9*-b-g}0\(1QNk36?)bK')(9 *K3(+?Z5K390ZD9N-Nk(qD9'0)3€JP(n?),4l65-,}Š[.qD95d0)g4AMD# ‹(+.+0)(I*‡JHA

D95dA€D9eT36b-5d0398CNk(I?@0),/.+,/*-(L#

0

10

20

30

40

50

60

70

80

−2

−1.5

−1

−0.5

0

0.5

Intrinsic Rate of Increase

Concentration Exposure

Intrinsic Rate of Increase (r

m

)

Neonate

Adult

R,4l6b-'@( 6# "F5H0)'),45-?),4.'@3 ih0)XS>DL0)(\8;36'!>(+365[DL0)(I?:D95-*S‚*-bKg402?

&.I36g436l6,/?02?"X[DqG6(‚?@02bK*-,/(+*x02XK(‚*-, ‹(+')(I5K.I(I?jJP(10Fio(+(I5Y,45-?@02D95d0fD95-(+36b-?j'2DL0)(>398‹,/5-.+')(ID9?)(†UMLˆ`

D95-*S02X-(,45H0)'),45-?),4.Z')DL02(398R,/5-.+')(ID9?)(=U o`…#‚.+.I36'@*-,/5-ln023 ‰ [0)X-($,45-?@02D95d0fD95K(I36b-?:'2D02(398

,/5K.I')(ID9?)(j,/?RD>eT(qD9?@b-')(j398-02XK(hD9J-,4g/,40FA‚398kD>Nk36N-bKg DL0),/365023>,/5-.+')(qDL?)((…QKNk365-(+5d02,DLg/g4A3 G9(I'R0),/eT(9#

hX-,4?j')DL02(:,/?rl6,4G9(I5xJHA ML

(

g45

(

(25)

?),4e†b-g/DL02,4365x,4?')bK5s#r,/5K.I(ij(\5-(I(+*D†?@,/5-l9g/(>GLD9g4b-(398 Ls023.+36exN-D9')(‚0)3 :Kij(\36JK0fDL,/5 36bK'

MLkGLD9g/b-(:JdAnŠ[5-*-,45-l$0)X-(>GLD9g4b-((IwHb-,4GLD9g4(I5d0r023$0)X-(‚Nk36N-bKg DL0),/365nl6'@3 ih0)X=')DL02(>365-.I(:02X-(:?@02D9J-g4(

D9l6(E*-,/?02'),4J-bK0),/365SX[D9?Jk(I(+5c')(qDL.fX-(I*ssiXK,/.fXm,/?&y@b-?@0*-(+0)(I')eT,/5K,/5-lT02X-(DL?@AOexN02390)(9# R,4l6b-')(

6#<*-(IN-,4.+0)?02X-(h,45-?@02D95d0fD95-(+36b-?"*-,4')(I.10"ex(ID9?)bK')(h398Pl6')3uih0)XT')DL02(ZUML;`C8;39'5-(I395[DL02(:D95K*xDL*-b-g40

(1QNk36?)bK')(‚Jk390)X D9?jDZNk36N-bKgDL0),/365TD95-*x*[D9,4g4A6#p‚?ri,40)XT0)X-( GLD9g4b-(I?+OD$5-(+ldDL02,}G6( MLk,/eTN-g/,4(I?

Nk36N-b-g/DL02,4365$(…QO02,/5K.+02,4365s#R3-9D9g40)X-36b-l6XED95D9?AOexNK0)3902(j,/?p5-390R')(qDL.fX-(I*†8;36'p5-(+365[DL0)((1QNk36?)b-'@(

398 6ŽN-N-eSKij(ED9'@(5-390?)bK')N-'@,/?)(+*JdAY0)X-(I?@($'@(I?)bKg402?+#

0

10

20

30

40

50

60

70

80

90

100

−2

−1.5

−1

−0.5

0

0.5

Average Instantaneous Growth Rate for Neonates

Time

Intrinsic Rate of Increase (r

i

)

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

0

10

20

30

40

50

60

70

80

90

100

−8

−7

−6

−5

−4

−3

−2

−1

0

1

Daily Instantaneous Growth Rate for Neonates

Time

Daily Intrinsic Rate of Increase

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

0

10

20

30

40

50

60

70

80

90

100

−0.1

−0.05

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

Average Instantaneous Growth Rate for Adult Exposure

Time

Intrinsic Rate of Increase (r

i

)

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

100 ppm

0

10

20

30

40

50

60

70

80

90

100

−0.4

−0.2

0

0.2

0.4

0.6

0.8

1

Daily Instantaneous Growth Rate for Adults Exposure

Time

Daily Intrinsic Rate of Increase

Control

10 ppm

20 ppm

40 ppm

60 ppm

80 ppm

100 ppm

(26)

hX-(\D9b02X-36'@?398 ‰ PX[DqG6(8;36b-5K*=02X[D0 ML.+36b-g4* JP(‚D9N-N-g4,/(+* D9?oDE?)b-JK?@02,}02bK0)(‚8;36' DL?

DNP39N-b-gD02,/395=(I5K*-NP39,/5d0ex(ID9?)bK')(IeT(I5d0I#rhXK(+A 8;36bK5-*€5-3?),4l65-,4Š-.qD95d0*-, ‹(+')(I5K.I(TU

6 `

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D9'@(Tg4,/?02(I*ci,40)Xm02X-(T.I36'@')(+?)Nk365-*-,45-l G9DLg/b-(+?Z,45BCD9J-g4( 6#:K#x>?$eT(I5d02,4365-(+*D9Jk3uG6(LC36bK'

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ML #€!>(+G9(I'02X-(+g/(I?@?I&(+.I36g436l6,/?02?ij36b-g4*g/,4_9(n023Sb-?)(T02X-( MLrGLD9g/b-(+?$,45-?@0)(qD9*B398o0)X-( G9DLg/b-(+?

Jk(I.qDLb-?)( ,4?he†b-.fXSex36'@(\*-,4tT.+b-g40h0)3T.ID9g/.+b-g D02(9#

CD9J-g4( 6#: &F5H0)'),45-?),4. E"(I')?@b-?:F5-?@02D95d0fD95-(+36b-?>>D02(398CF5-.I'@(qD9?@(

> ?&(?1 '!2# " ? , ?9# * + 49

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ML ML

o365d02'@36g # #dŒ # #dŒ

qŽ9N-N-e #aŒ6Œ # #dŒ6Œ #Œ9Ž

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uŽ9Ž9N-N-e #Ž3: #aŽ6Œ

‰ ,40)Xx36bK' GLD9g4b-(9H36'&0)X-(I36'@(+0),/.qDLg/g4AE36b-' ML^GLD9g/b-(Ldio(>.qD95x.qD9g4.Ib-g/DL02(h36bK'rŠ[5-D9g^N[D9'2DLen{

(+0)(I'@?h398„,/5d02(+')(+?@0q# \'2D9NKX-?3980)X-(,/5d0)'),/5K?),/.>J-,/'02XYDL5-*Y*-(qD02XY'2D02(I?8;39'o(qDL.fXS.+365-.+(I5d02')DL02,4365

(27)

*-(+.I')(ID9?)(SiDL?=e†b-.fX ex39')(Y.I395-?),4*-(I')D9J-g/(Y8;36'n5-(+365[DL0)((…QNP36?@b-')(L# (qDL0)X ')DL02(€,45-.I'@(qD9?@(I*

D9?.I395-.I(+5H0)'2DL0),/365€398CNk(I?@0),/.+,/*-(\,45-.I'@(qD9?@(I*8;39'5-(I395[DL02((…QKNk36?@b-')(LkXK3 ij(+G9(I'+‹D95S,/5K.I')(ID9?)(,45

Nk(I?@0),/.+,/*-(\.+365-.I(+5d02'2D02,/395*-,4*€5-390?)(+(Ie0)3=D# P(I.10h02X-(*-(IDL02XS'2D02(\8;36'D9*-bKg40(1QNk36?)bK')(9#

0

10

20

30

40

50

60

70

80

0

0.05

0.1

0.15

0.2

0.25

0.3

0.35

0.4

Intrinsic Birth Rate

Concentration Exposure

Birth Rate (b)

Neonate

Adult

0

10

20

30

40

50

60

70

80

0

0.2

0.4

0.6

0.8

1

1.2

1.4

1.6

1.8

Intrinsic Death Rate

Concentration Exposure

Death Rate (dr)

Neonate

Adult

(28)

*

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(29)

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(30)

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(31)

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ij36')_H,/5Kli,}02XTD.I395H0),/5Hb-36bK?reT3O*-(IgOiX-(I5Tb-?),45-l$-P9ij(:D9')(DLJ-g/(0)3\*-(+0)(I')eT,/5K(hNP36NKb-g DL0),/365

*KAO5[D9eT,/.+?h5-390>DqGLD9,/g/D9J-g/(:i,40)X(+?)g/,4(9#

(40)

0

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Linear Interpolation of Adult Recruiment for 60ppm

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Density

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Linear Interpolation of Adult Recruiment for 80ppm

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Density

(41)

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Linear Interpolation of Adult Recruiment for the Control

Age

Density

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Linear Interpolation of Adult Recruiment for 10ppm

Age

Density

R,4l6b-')(>Œ# C‚*-b-g}0j:(+.I')bK,402eT(I5d0jD95-*=%C36NKb-gDL0),/365xO0)')b-.102b-'@(:8;39'r0)X-(o395H0)')36gkD95K* uŽN-N-e

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(42)

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Linear Interpolation of Adult Recruiment for 20ppm

Age

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Linear Interpolation of Adult Recruiment for 40ppm

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(43)

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Linear Interpolation of Adult Recruiment for 60ppm

Age

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Linear Interpolation of Adult Recruiment for 80ppm

Age

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(44)

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References

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