Memoirsofthe MuseumofVictoria53(1): 115-123 (1992)
THYSANOPTERA (INSECTA) FROM THE 1985 AND 1986 ZOOLOGICALEXPEDITIONS TO THEKRAKATAUS, INDONESIA
By Richard zur Strassen
ForschungsinstitutSenckenberg. Senckenberg-Anlage25, D-6000Frankfurtam Main, Germany
Abstract
zurStrassen, R., 1992.Thysanoptera (Insecta)from the 1985and 1986Zoological Expe- ditions to the Krakataus, Indonesia. MemoirsoftheMuseumofVictoria 53: 115-123.
TheThysanopteracollectionsoftheLa TrobeUniversity/LlPl 1985and 1986Zoological ExpeditionstotheKrakatauislandsintheSundaStrait,Indonesia, include26species.Two
species are described asnewandfigured:Mymarothripsbicolorsp. nov.,andApollolhrips karnyisp. nov.A furtherfive speciesarerecorded from Indonesiaforthe firsttime:Ani-
sopilothrips venustulus(Priesner),Apollolhripsbhattii Wilson, Panchaetothrips holtmanni Wilson, Plectrothrips eximius Ananthakrishnan, and Strepterothrips orientalis Anantha- krishnan.
Introduction
The 1985and 1986Zoological Expeditionsto the Krakataus in the Sunda Strait, Indonesia, during August (1985) and September (1986), werecarriedout byzoologistsfrom Indonesian andAustralian institutions. Ageneral introduc- tion tothe 1984and 1985 expeditionswaspro- vided by Thornton and Rosengren (1988). A
report on the Thysanoptera collected on the 1984 expedition was presentedby zurStrassen (1991).
About two dozen samples ofthrips (Thysa- noptera) were collected in 1985, and four samplesin1986.One1984 sampleisincludedin the present paper.
Thethysanopteranswereobtained mainlyby beatingand sweeping vegetation, and in a few cases by Malaise traps. Many specimens are damaged, severalofthem havetheantennaeor parts of them broken off. Therefore, not all
specimens have been identified tospecies.
Material is deposited in the Lembaga Ilmu Pengetahuan Indonesia (LIPI), the Forschung- sinstitut Senckenberg at Frankfurt am Main, Germany (SMF) and the Museum ofVictoria,
Melbourne, Australia (NMV).
The species dealt with below are arranged systematically under families. Within thesethe genera are arranged alphabetically as are the speciesunder each genus.
Aeolothripidae
Mymarothrips bicolorsp. nov.
Figure 1
Material examined. Holotype: Krakataus.PanjangI.,
by sweeping (235-D2B), 17 Aug 1985, LIPI (female).
Paratype:Krakataus, SertungI.,bybeating twigsin forest(244-CS), 18 Aug 1985, SMF(1 female).
Diagnosis. Head about as long as broad. Body
2-coloured, anterior part of body largely pale yellow,abdominalsegmentsV-Xbrowntodark
brown, margins of head darkened; forewings whitein proximalhalf,brownin distalhalfwith large circular colourless apical patch; dorsal setae on abdominal segments IX and X
yellowish.
Description(female macropterous). Totallength (distended) 1470-1550 urn. Head between the eyes yellow, anteriormargin andcheeksasdeep as eyeswidth brown,mouthconeyellow;anten- nae dark brown, segmentsIand VII-IX a trifle paler. Pronotum yellow with lateral margins brownish, setae pale. Pterothorax and abdomi- nalsegmentsI
—
IIIpale yellow,segment IIIwith small pale brown spot near lateral marginjust behind subbasaltransverseline,segment IVpale
brownmottledposteriorly orlaterally,otherwise yellow, segments V-VII coloured as in IV, or uniformly brown, segment VIII somewhat darker than VII or even darkbrownasabdomi- nal segments IX and X; setae on latter 2 seg-
ments yellowish. Legs pale yellow. Forewings
(Fig. 1) white in proximal halfincluding veins, setaeandscapus, brownin distalhalfexceptfor large circular white patch almost at apex, vein setae in this part dark brown; hindwings simi- larlycoloured, but thebrownofa slightly paler tinge and the white patch not as clearly mar- ginated as in forewing.
15
https://doi.org/10.24199/j.mmv.1992.53.05 30 May 1992
116 R. ZURSTRASSEN
200pm
FigureI.Mymarothripsbicolorsp.nov.,holotype female,LIPI(right forewing showingthecolour pattern as seen by meansofphasecontrast. Drawnby Andrea Vesmanis.
Headaboutaslongasbroad, ante-ocularseta dark brown, 38-41 urn long, other head setae pale. Length in urn of antennal segments of holotype: I 31, II 49,III 84, IV64,V 59,VI 54, VII 57, VIII 19, IX 17. Pronotum 139-154 urn long, 178-205 um broad, subanteromarginal setaeaslongasorsomewhat longer(57-69um) thananteromarginals(51-56 um), posteroangu-
larsetae 74-86 um long. Pterothorax 300-318
um long, 270-298 um broad; hind tibia 218- 230um,hindtarsus103-115umlong.Forewing 820-880 um long, across middle 107-115 um
broad,beforeapex 192-215p broad, distance betweenanteriorandposterior veinatcrossvein 17-23 um, apicalsetae ofposteriorvein 63-75
um long. Abdominal segments IX+X together 225-240umlong, widthacrossbase ofsegment IX 146-161 um; segment IX with seta S, 126-
129um,S2 138-149um,dorsalseta86umlong, segmentX withsetaS, 115-121 um long. Ster- niteVIwith4-6,sterniteVIIwith 10-13access- orysetae. Ovipositor355-375 um long.
Discussion.ThenewspeciesM. bicolorisreadily distinguishedfrom itscongeners by thecolour- ation. The type species, M. ritchianus Bagnall (1928)fromtheEthiopianregion, isagenerally pale species with orange and red internal pig- mentation; pterothoraxat lateralmargin,meso- and metasternum,abdominalsegmentsI,distal halfofIX andall ofXareyellowishbrown;the forewing is very pale brownish at its base, the thirdand fourthfifthhasa slightlydarkertinge at the level ofthe cross vein between anterior andposteriorvein,andacross thebroadestpart ofthewing. The twospecies from the Oriental region, M. bolus Bhatti, 1967 and M. garuda
Ramakrishna and Margabandhu, 1931 are
muchdarker thanthenewspeciesand havedark setaeallalongtheveinsoftheforewing, the costa ofwhich isdark in itstotal length.
Thripidae
Anisopilothrips venustulus (Priesner) Heliolhripsvenustulus Priesner, 1923: 89.
Anisopilothripsvenustulus. —Stannard and Mitri.
1962: 187.
Material examined. Krakataus: PanjangI.,by sweep- ing(164-N), Sep 1984,LIPI (1 female).
Distribution.Perhaps of Caribbeanorigin,intro-
duced tomany subtropical andtropical islands and several countries nearthe sea; foliicolous.
Not previously recorded from Indonesia.
Apollothrips bhattii Wilson
ApollothripsbhatliiWilson, 1972: 52.
Material examined. Krakataus: PanjangI.,by sweep- ing(164-N),Sep 1984, LIPI(1 female);PanjangI.,by sweeping(330-CE), 25Sep 1986,SMF(1 male). Ser- tung I., Spit, by beating in the transition zone (244-HM),NMV(1 female),andby sweepinginforest (245-L2C), both 18Aug1985. LIPI,SMF(2females);
Panjang I., South Ridge, by sweeping in forest (340-DD), 27 Sep 1986, NMV(1 female).
Distribution. Until now known only from the type series from Madhya Pradesh, India; per- haps living on ferns.
Apollothrips kamyisp. nov.
Figures 2-4
Material examined. Holotype: Krakataus, SertungI..
THYSANOPTERAFROMTHE KRAKATAUS 117
100pm
Figures 2-4. Apollothripskarnyisp.nov.,holotype female(LIPI).2,antennal segmentVIdorsally;3,meso- and metanotum; 4, tergiteVI. Drawn byAndreaVesmanis.
THYSANOPTERAFROMTHE KRAKATAUS 119
Materialexamined. Krakataus: RakataI.,OwlBay,by sweepingfromlowcoastalvegetation(225-JI),25Aug
1985, LIPI (1 male). Anak Krakatau I., West shore, from Malaise trap(313-EM), 24 Sep 1986, NMV (1 male). SertungI.,South Ridge,by sweepingin forest (340-DD), 27 Sep 1986, LIPI, SMF(2 males).
Distribution. Japan, Taiwan, Sumatra, Java,
Vanuatu (= New Hebrides), Tonga; florico- lous.
Panehaetothrips holtmanni Wilson PanchaetothripshohmanniWilson, 1975: 186.
Material examined. Krakataus: PanjangI., by sweep- ing low vegetation (235-EU), 16 Aug 1985,"LIPI (1 female);PanjangI.,NEshore,by sweeping (235-BH),
17Aug1985,SMF( 1 female).SertungI.,SouthRidge, by sweepingin forest(340-DD), 27 Sep 1986, NMV
(1 female).
Distribution. Until now known only from the type material from NewGuinea; foliicolous.
Remarks. The three specimens are regarded as belongingtoP. holtmannibecause oftheforked senseconesonantennalsegmentsIIIand IV(not mentioned in the original description because only segments I and II are present in the type specimens); the weak and pale seta S, on hind marginoftergiteIIwhichisabouthalf aslongas setaS2 (not asstrongand longas);the subbasal (antecostal) line on tergites II—VIII with many
posteriorlydirected small notches; and the dis- cal campaniform sensilla on metanotum being separatedbymore than twooftheirowndiam-
eters. However, innone ofthe specimens dealt withisthere aminute setamidwaybetween the posterior ocelli and the eyes as has been described forthe type specimens.
Scirtothrips dorsalis Hood
Scirtothripsdorsalis Hood. 1919: 90.
Material examined. Krakataus: Anak Krakatau I.,
from Malaisetrapatouterconc(2l3-SJ),18Aug1985, LIPI(1 female);AnakKrakatauI., fromMalaisetrap
NE foreland(213-PK), 19Aug 1985,SMF(1 male).
Distribution. From Japan, New Guinea, the Solomonsand Australia, westwardto Pakistan, also S. Africa (perhaps introduced); florico- lous/foliicolous.
Selenothrips rubrocinctus (Giard) Physopus rubmcincta Giard. 1901: 264.
Selenothrips rubroeinetus.—Hood, 1913: 150.
Material examined. Krakataus: AnakKrakatauI.,by beating broad leaved plants (214-FD), LIPI, SMF,
NMV(7females),andby anotherbeatingbroadleaved plants (214-GG),21 Aug 1985,LIPI (1 female).
Distribution. Circum-subtropical and -tropical;
foliicolous.
Thrips hawaiiensis (Morgan)
Euthrips hawaiiensisMorgan, 1913: 3.
Thrips hawaiiensis. — Priesner. 1934: 266.
Material examined. Krakataus: Anak Krakatau 1.,
from Malaise trap on outer cone (213-SJ), 18 Aug
1985, LIPK1 female).
Distribution. Southern States ofUSA, Pacific, Japan, Oriental Region; floricolous, polypha- gous.
Zonothrips karnyi Priesner ZonothripskarnyiPriesner, 1926: 260.
Material examined. Krakataus: AnakKrakatau 1., by sweeping(215-EL),22 Aug 1985(LIPI), 1 female.
Distribution. Indonesia (Java); recently recorded from Rakata I. (zur Strassen, 1991).
Phlaeothripidae
Ecacanthothrips tibialis (Ashmead)
Idolothrips tibialisAshmead, 1905: 20.
Ecacanthothrips tibialis. — Palmer and Mound, 1978: 156.
Material examined. Krakataus: PanjangI.,by sweep- ing low vegetation (235-EU), 16 Aug 1985, LIPI (2 females).RakataI.,OwlBay,by sweepinglowcoastal vegetation(225-JI), 25Aug 1985. NMV(1 female).
Distribution. Indo-Australian Region, Japan, Mauritius, Rodrigues I., Tanganyika; fungivor- ous.
Elaphrothrips curvipes Priesner Elaphrothrips curvipes Priesner, 1929: 206.
Material examined. Krakataus: Sertung I., Spit, by beating in Casuarina forest (244-AN), LIPI (1 male and3 females),samelocality,bybeatinga Casuarina tree(244-IO),NMV( 1 male),andby sweeping under- story in Casuarina forest(245-AH), LIPI (1 female),
all 18Aug 1985.Anak KrakatauI., by beatingbroad leaved plants (214-FD), LIPI. SMF (2 males and 1
female),andby another beatingbroad leaved plants (214-GG), NMV (I male and 1 female), all 21 Aug
1985.
Distribution. Southeast Asia inclusive of Indo- nesia; fungivorous.
118 R.ZUR STRASSEN
South Ridge,by sweepinginforest(340-DD), 27Sep 1986, LIPI (female).
Diagnosis. Mouthcone 1.7timesas long aspron- otum; discal setae on pronotum thin, not thickened; mesonotum in median third with sculptureofanastomosing transverse lines; ter- gites II—VIIsmooth inmedian part betweenthe 2 setae S2, no transverse lines around area of setaeS,,discalcampaniform sensillaontergites IV-VIII situated further cephalad than setae S,.
Description(female macropterous).Total length (distended) 1520 urn. Head, pro-and ptcrotho- rax darkgreyish brown, metanotumalong mid- linewithnarrowlongitudinal palerarea, tergites medially and sternitesgreybrown, tergites lad- ing laterad to almost white. Antennae dark brown, segments IV-VI immediately behind theirbasalmargin each with narrow, transverse whitish band. Legs paleyellow;wings brownor greyishbrown. Body setaepalebrown, thoseon abdominal segments IX and X dark brown.
Head about 1.4 times as broad (160 um) as long (115 urn). Dorsal surface densely sculp- tured withanastomosing transverse lines, setae short, not longer than 18 um. Mouth cone almost pointed, 220 um long, surpassing hind margin of pronotum, maxillary palpi 69 um
long.Antennaeabout410umlong(lengthofseg-
ments II and III in the unique specimen not exactlytomeasure), segments 111 and IVbottle- shaped, each with stout, forkedsense cone with 57-63 um long branches. Innerandouter sense cone on segment VI (Fig. 2) arising from long linear base, thatofthe innercone 17 um, thatof theoutercone 29umlong,thefree distal partof the innercone31 um,that oftheoutercone 26
Um long.
Pronotum 155um long, 161 umbroad across anterior margin, 212 um broad across widest part;surface withoftenanastomosingtransverse linesofwhich40cross the longitudinal midline, and with about 60 short discal setae which are not thickened;hindmarginwithoutpronounced
setae. Pterothorax 298 um long,270 um broad.
Mesonotum(Fig. 3)initsmedianpartwithsame type ofsculpture as on pronotum; metanotum without campaniform sensilla. Hind tibia 200
umlong;forewingsslender,875um long,98um
broad across scapus, 38 um across middle,
anterior veinwith 3+6(-7) subbasalsetaeand 2 distal setae, latterverycloseto tip ofwing, pos- terior vein with 20 setae.
Abdominal tergites II—VII smooth in whole areabetweenthe 2 setaeS2(see Fig.4), noshort
transverselinesaroundsetaeS,; discalcampan- iformsensillaontergitesIV-VIIsituated further cephalad than setae S,; lateral parts oftergites with 12-20 setae laterad of seta S2; ctenidia wanting. Abdominal segments IX+X together
161 pmlong,dorsallywithoutsculpture,setaS,
on IX69 um, S275 um, dorsal seta55 um long, seta S, on X 92"urn, S, 77 um long. Ovipositor 260 um long.
Etvmologv.Thespeciesisdedicatedto Prof.Dr.
H.'Karny(1886-1939), the famous entomolog-
ist who for decades studied the Indonesian Thysanoptcra.
Discussion. The new species A. kamyi is separ- ated from A. bhallii Wilson, 1972, hitherto the only known species in the genus, by the longer
mouthcone (220pm;in.4. bhallii 140- 160pm);
bythenormal shape ofthe discalsetaeon pron- otum (not being stout and thickened): by the different sculptureofthemesonotumwhere the transverselinesarc oftenanastomosing(instead of hardly anastomosing and running very closely); by the smooth median part oftergites II-VII between the setae S[ (instead of short transverse lines around the setae S,); by the location ofthe discal campaniform sensilla on
tergites IV-VIII,situatedfurthercephalad than setaeS, (insteadoffurthercaudad,asin.A.bhal-
lii).
Astrothrips globieeps (Karny) Heliotkrips gtobicepsKarny, 1913a: 125.
Astfothfipsglobiceps. — Karny. 1923: 331.
Material examined. Krakataus: Sertung I.. South Ridge,by sweeping in forest (340-DD). 27 Sep 1986.
LIPI(1 female).
Distribution. MelanesiatosoutheastAsia: folii- colous. Just recently recorded from Sertung I.
and Rakata I. (zur Strassen, 1991).
Dorcadothrips trifasciatus(Priesner) Taeniothripstrifasciatus Priesner, 1936: 323.
Dorcadothripstrifasciatus. —Bhatti, 1978: 169.
Material examined. Krakataus: Rakata I.. Owl Bay.
from Malaise trap (223-AJ). 27 Aug 1985 (LIPI), 1
female.
Distribution. This is the first record of the species sinceitsdescriptionfrom Sumatra.Wai Lima (Lampongs); perhaps foliicolous.
Megalurothrips formosae (Moulton)
Taeniothripsformosae Moulton, 1928: 298.
Megalurothrips formosae. — Bhatti, 1969: 241.
120 R. ZUR STRASSEN
Ethirothrips stenomelas (F. Walker) Phbeothrips stenomelas Walker, 1859: 224.
Ethirothrips stenomelas. — Mound and Ralmer.
1983: 57.
Materialexamined. Krakataus: RanjangI., by sweep- ing low vegetation (235-EU), 16 Aug 1985, LIPI (I
female).
Distribution. Pacific, Indo-Australian Region, India, Madagascar; sporophagous.
Haplothripscertus Priesncr Haploihripscertus Pricsner, 1929: 194.
Material examined. Krakataus: Anak Krakatau I.,
from Malaise trap on outer cone (213-S.I), 18 Aug
1985, LIRI (I male).
Distribution. New Guinea. S. China, Taiwan, Sumatra, Mentawei Archipelago; on Cypcra- ccae.
Haplothripsganglbaueri Schmutz
Haplothripsganglbaueri Schmutz, 1913: 1034.
Material examined. Krakataus: Anak Krakatau I.,bv beating grasses (214-EF). 21 Aug 1985. LIPI (I
female).
Distribution. West Pacific, Oriental Region;
graminicolous.
Holurothrips ornatus Bagnall llolurothripsornatus Bagnall, 1914b: 376.
Materialexamined. Krakataus: Panjang I.,bvsweep- ing low vegetation (235-EU), 16 Aug 1985,"LIPI (1
female).
Distribution. Oriental Region; sporophagous.
Machatothrips antennatus (Bagnall) Adiaphorothrips antennatusBagnall. 1915: 594.
Machatothripsantennatus. —Mound, 1968: 133.
Material examined. Krakataus: RakataI., S.Face,by beating(224-GL). 25 Aug 1985 (LIPI), I male.
Distribution. S. China, Malaya, Singapore, W.
Sarawak, Indonesia; sporophagous.
Meiothrips menoni Ananthakrishnan MeiothripsmenoniAnanthakrishnan, 1964b: 99.
Material examined. Krakataus:PanjangI., bvsweep- ing low vegetation (235-EU), 16 Aug 1985,"LIPI (1
male).
Distribution. Oriental Region; sporophagous.
Nesothrips brevicollis (Bagnall) Oedemothripsbrevicollis Bagnall, 1914a: 29.
Nesothripsbrevicollis.— Mound, 1968: 140.
Material examined. Krakataus: PanjangI.,by sweep- ing(
I64-N),Sep1984,LIPI(1 female-brachypterous).
Sertung I.,Spit, by beating in Casuarina forest(244- AN). 18Aug1985, LIPI,SMF,NMV(1 male-macrop-
tcrous. 1 fcmale-macropterous and 1 female- brachypterous);sameisland,forest,by sweeping(245- L2C), 19 Aug 1985, SMF (I male-hemimacropte-
rous).AnakKrakatau I.,bybeatingfrom broadleaved plants(214-FD). 21 Aug 1985, LIPI,NMV(1 fcmale- macropterousand 1 female-brachypterous).
Distribution. Old World tropicsand subtropics;
sporophagous.
Nesothrips lativentris (Karny) Rltaehothripslativentris Karny, 1913b: 129.
Nesothripslativentris.— Poundand Palmer, 1983- 48.
Material examined. Krakataus: Sertung I., Spit, by beatinginthetransitionzone(244-BP), 18Aug 1985, LIPI (I malc-brachypterous). Anak Krakatau I., by beatingbroad leaved plants(214-GG), 21 Aug 1985, LIPI. NMV (1 female-macropterous and 1 female- brachypterous).
Distribution. Old andNewWorldtropics;spor- ophagous.
Plectrothrips eximius Ananthakrishnan
Pleelrathrips eximius Ananthakrishnan, 1969 296.
Material examined. Krakataus: Rakata I., Owl Bay from Malaise trap (223-AJ). 27 Aug 1985, LIPI (1 female). Anak Krakatau I., from Malaisetrap (313- CN). 24 Sep 1986. SMF(1 female).
Distribution. Described from a unique female from Madras, S. India; fungivorous. Okajima (1981:319) gave anadditionalrecordfrom Sin- gapore.
Remarks. The two specimens from the Kraka- taus differ to some extent from the type speci-
men but nevertheless are regardedas "belonging toP. eximius.Thebodycolourisuniformdark, there arenoyellowtinged portions,eventhelegs aredark brown, only thedistalpartsofthetibia are somewhat paler, the tube has a tinge of red.
Pygothripsvicinus Okajima
PygothrfpsvtcinusOkajima, 1990: 97.
Material examined. Krakataus: Sertung I., Spit, by
THYSANOPTERAFROMTHE KRAKATAUS 121
beating in Casuarinaforest (244-AN), 18Aug 1985 LIPI. SMF,NMV(6females).
Distribution. Ryukyu Islands (Japan), Java, Thailand; sporophagous.
Remarks. The epimeral suture is complete in these six females whereas it is described as incomplete inthe typematerial.
Strepterothripsorientalis Ananthakrishnan
StrepterothripsorientalisAnanthakrishnan, 1964a- 118.
Material examined. Krakataus: Anak Krakatau I.,
from Malaise trap (313-CN). 24 Sep 1986, LIPI (1 female-macropterous).
Distribution. Hawaii,Fiji,Taiwan,Thailand, S.
India. Not previously recorded from Indo- nesia.
Streptothrips mirabilisPriesner Streptothrips mirabilis Priesner, 1932: 58.
Material examined. Krakataus: Sertung I., Spit, by beating in Casuarina forest(244-AN), 18 Aug 1985, LIPI (1 female).
Distribution. This is the first record of the species sinceits description fromSumatra,Wai Lima(Lampongs).
Discussion
Inafirstreport (zur Strassen, 1991)onThysa- noptera recently collected from the Krakataus 25 species were recorded from the 1984 Expedition to this island group in the Sunda
Strait. The second report on newly submitted material dealswith 26 species mostly takenon the 1985Expeditiontothe archipelago.The new
material contains a further 15 species not pre- viously recorded from the Krakataus; eight of these species are also additions to the known
fauna of Indonesia, of which two species are evennewly described(Mymarothripsbicolorsp.
nov., Apollothripskarnyisp. nov.). Twospecies are recorded here for the first time since they were described long ago, i.e. Dorcadolhrips tri-
fasciatus Priesner, 1936 andStreptothripsmira-
bilisPriesner, 1932.
Fourteen(58%)ofthe species are represented by only a single specimen. The corresponding percentage forthe 1984 material was 36%.
Acknowledgements
The author is grateful to Professor I.W.B.
Thornton, Head ofthe Departmentof Zoology,
La Trobe University at Bundoora, Victoria, Australia, and to Mr P.A. Vaughan. Research AssistantontheKrakatauExpeditions,formak-
ingthismaterial available. Thanksareduealso to the participantsofthe 1985 and 1986 Expe- ditionsfor collecting thespecimens, and tothe
many sponsors forhaving rendered the project possible.
References
Ananthakrishnan, T.N., 1964a. Thysanopterologica Indica—I.EntomotogiskTidskrifi85(1-2):99- 120.
Ananthakrishnan,T.N., 1964b.Acontributiontoour knowledge oftheTubulifera(Thysanoptera)from India.Opuscula entomologicaSupplementum25:
1-120.
Ananthakrishnan, T.N., 1969. Mycophagous Thysa- noptera— II. OrientalInsects 3(3): 289-299.
Bagnall,R.S.,1914a. Brief descriptions ofnewThysa- noptera - II. Annals and Magazine of Natural History(8) 13: 22-31.
Bagnall,R.S., 1914b. Brief descriptionsofnewThysa- noptera - IV. Annals and Magazineof Natural History(8) 14: 375-381.
Bagnall, R.S., 1915. BriefdescriptionsofnewThysa- noptera - VI. Annalsand Magazine ofnatural History(8) 15: 588-597.
Bagnall.R.S..1928.Preliminary description ofMyma-
rothrips ritchianus,anewtypeof Thysanopteron.
Annalsand Magazineof NaturalHistory (10) 1:
304-307.
Bhatti,J.S., 1967. ThysanopteraNova Indica. pp 23.
Published privately by the author. Delhi, 1 1
March 1967.
Bhatti,J.S., 1969. ThetaxonomicstatusofMegalitro- thripsBagnall (Thysanoptera: Thripidae). Orien- talInsects 3(3): 239-244.
Bhatti, J.S., 1978. Apreliminary revision of Taenio- thrips(Thysanoptera:Thripidae).OrientalInsects
12 (2): 157-199.
Giard,A., 1901. Surun Thrips (Physopusrubrocincta nov.sp.)nuisibleau Cacaoyer(Thys.).Bulletinde la Societe Entomologique de France 1901 (15):
263-265.
Hood, J.D., 1913. On a collection ofThysanoptera from Porto Rico. InsecutorInscitiaeMenstruus 1
(12): 149-154.
Hood, J.D., 1919. On some newThysanoptera from southern India. Insecutor Inscitiae Menstruus 7 (4-6): 90-103.
Karny, H.. 1913a. Beitrag zurThysanopteren-Fauna von Neu-Guinea undNeu-Britannien.Archivfiir
Naturgeschichte (A)79(1): 123-136.
Karny, H., 1913b. H. Sauter's Formosa-Ausbeute:
Thysanoptera. Sapplemenia Entomoloi>ica 2:
127-134.
Karny,H., 1923. Beitrage zurmalayischenThysanop- terenfauna. Treubia 3 (3-4):277-380.
Morgan,A.C., 1913.NewgeneraandspeciesofThysa- noptera, with notes on distribution and food
