What is evolution?
The central and unifying theme of biology
The term “evolution” means gradual change over time
Geologic evolution: the slow, continuous change of the earth itself
Evolution
is the process by which
one species gives rise to another.
Confusion sometimes arises as to whether Evolution is a theory or a fact.
The theory of Evolution deals with how Evolution happens. Theories in science explain phenomena.
Evolution is also a fact as there is a huge amount of
Misconceptions about
Evolution
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Pope Francis has said:
“When we read the creation story in Genesis we run the risk ofimagining that God was a magician, with a magic wand which is able to do everything. But it is not so. He created beings and let them develop according to internal laws which He gave every one, so they would
develop, so they would reach maturity.”
Fossil: a trace or remains of an organism that has been preserved by natural processes
Mostly the hard parts (bones and shells) are preserved
Fossils allow paleontologists to date the organisms The layers of sediment in which the fossils are
formed can give some idea of relative dating
Problems with Fossil
Records
Fossils rarely form; most organisms either decay completely or are consumed
Fossils that do form rarely survive long enough for us to find them
Fossils are rarely accessible – they are buried too deep for us to find
2. Comparative
Anatomy
Comparative Anatomy refers to the
structural similarities and differences
between living things.
Similar structures in different organisms
points to similar ancestry. The more
Homologous Structures
Parts of different organisms that have
similar structures, but different forms or
functions.
Analogous Structures
Different organisms have organs with the same function, but different underlying structure.
They are thought to be evidence for evolution along two different lines.
Vestigial Structures
FYI…
Humans have over 100 vestigial structures!
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Coccyx (tailbone)
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Appendix
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Wisdom Teeth
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Goosebumps
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Infant grip reflex
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Muscles to move nose and ears
Embryological
Similarities
Closely-related species have similar patterns
of development
In embryonic stages, homologous structures may appear that may not appear in the adult forms of two organisms
For example, humans have a very early stage in utero with gill-slits. This points to a common ancestor with fish.
Humans also have a tail for much of their
Molecular Similarities
Due to genome sequencing, we can now examine similarities in DNA
Because every organism inherits DNA from
their ancestors, the DNA of every living
comes originally from the earliest forms of
life
Cytochrome c
One protein, which is needed for cellular
respiration, is called Cytochrome c.
Because of mutations in DNA,
Differences among cytochrome c are more
pronounced the further back a common
ancestor between two species is (the longer
ago the species diverged)
Cytochrome c
comparisons
Humans and chimps – 1 amino acid differs
Humans and pigs – 10 amino acids differ
Jean Baptiste Lamarck
Presented one of the first theories of evolution in 1809Was convinced that species were not constant
Thought that changes in
First Principle: Law of
Use and Disuse
◦
The more an animal uses a specific part of
their body, the stronger and
better-developed that part becomes.
◦
The less a part is used, the weaker and
Second Principle:
Inheritance of Acquired
Characteristics
Characteristics that an organism acquires
Classic Lamarck
Example
Giraffe’s long necks:
◦ Lamarck knew the ancestors of modern giraffes
had short necks
◦ He thought they would have run out of grasses
and low leaves to eat and had to stretch their necks to get food
◦ Over time, the stretching resulted in their necks
getting longer
Charles Darwin
Son of a physician who wanted him to study medicine
Went on a voyage on the HMS Beagle in 1831, which lasted 5 years. He returned convinced that organisms evolve.
Made many observations of plants, animals, and insects
Made his most significant
Darwin’s Book
On the Origin of Species by Means of Natural Selection
Darwin’s Influences
Charles Lyell: The Principles of Geology. This proposed the earth was very old and changing over time (Geological Evolution)
Thomas Malthus: An Essay on the Principle of Population. This proposed that human food supply could not keep up with the growing population, leading to death by disease and starvation.
Darwin’s Theory of
Natural Selection
Read the text pages 420-421
Complete the worksheet
For the examples use the giraffe. How would
Darwin’s theory of evolution by natural
Hugo Marie de Vries
1848-1935
Dutch botanist and geneticist
Rediscovered the Laws of Heredity in 1890, without having read
Mendel’s work
Mutation Theory
Introduced the term “mutation” to describe
suddenly-appearing varieties of plants
Argued that new species are not formed by
continuous variations, but by sudden
Alfred Wegener
1912
Alfred Wegener proposed the theory of
continental drift – the idea that Earth’s
continents move. Despite publishing a large
body of compelling fossil and rock evidence
for his theory, it was rejected by most other
scientists. It was only in the 1960s that
August Weismann
1834-1914
Studied chromosomes
Was not convinced of
Lamarck’s theories and did experiments to disprove them
Weismann’s work formed the basis for Modern Synthetic Theory and provided an important link between genetic material and natural selection.
Evolution Heredity
Lamarck (Use + Disuse)Mendel (Inheritance)
Darwin (Natural Selection) de Vries (Mutations)
Weismann
Theodosius Dobzhansky
1937
Author of Genetics and the Origin of Species, a book that offers an explanation for how species actually came into existence.
Mutations crop up naturally all the time. Some
Modern Synthesis
Dobzhansky's ability to combine genetics and natural
history attracted many other biologists to join him in the effort to find a unified explanation of how evolution
happens. Their combined work, known as "The Modern Synthesis," brought together genetics, paleontology,
E. O. Wilson
1975
SOCIOBIOLOGY: THE NEW SYNTHESIS
Wilson used sociobiology and evolutionary principles to explain the behavior of social insects. He argued that all animal behavior,
including that of humans, is the product of heredity,
environmental stimuli, and past experiences, and that free will is an illusion.
The sociobiological view is that all animal social
behavior is governed by
Microevolution vs.
Macroevolution
Microevolution: the evolution of a
species as the environment changes.
Microevolution: Dogs
are Wolves!
The dog is example of how selection can change the
frequency of alleles in a population.
Dogs have been artificially
selected for certain characteristics for many years, and different
breeds have different alleles.
All breeds of dog belong to the same species, Canis lupus (the wolf) so this is an example of
Macroevolution:
Darwin’s Finches
However, if two populations of a species become isolated from
one another for tens of thousands of years, genetic difference may become marked.
If the two populations can no-longer interbreed, new species are born. This is called Macroevolution.
Darwin’s Galapagos finches are
Speciation
Species:
Group of organisms that are structurally similar and naturally interbreed to pass similarity on to offspring
Speciation: Refers to the formation of a new species.
Reproductive Isolation
Is what keeps members of one population
from interbreeding with members of
Barriers to reproduction and
recombination between species:
Pre-mating Barriers: Prevent mating from
occurring between two organisms of
different species.
Geographical Isolation
Divided by a natural barrier
◦ Ocean, mountain, desert, river…
◦ e.g. Until very recently polar bears and grizzlies
Ecological Isolation
Animals inhabit different habitats
◦ For example, white-crowned sparrows live in
Temporal Isolation
Different species breed at different times of
the year.
Behavioural Isolation
Courtship and mating rituals differ, so that members of different species won’t
interbreed.
◦ E.g. Female songbirds
only respond to the song of a male of the same species.
◦ Blue-footed booby dan
Mechanical Isolation
Reproductive structures between two species are incompatible.
Gametic Incompatibility
Sperm of one species cannot fertilize
egg of another
Hybrid Inviability
The hybrid (result of mating between
different species) does not survive to
adulthood.
E.g. It may show behavioural traits of both
parents, which are incompatible with each
other
Hybrid Infertility
Species which are similar can often reproduce. If, however, they have a different chromosome
number, this poses problems.
With unequal numbers of chromosomes
Following isolation…
Isolating a group of organisms separates its gene pool from the rest of the species.
Following that, mutation, genetic recombination, and natural selection can cause a different gene pool to evolve .
Adaptive Radiation
Refers to species spreading or radiating out
into different environments.
Each species will adapt to the pressures in its
own environment.
Other Patterns of
Evolution
Coevolution
The Rate of
Speciation
Gradualism
According to Darwin’s theory, new species arise
through gradual accumulations of small variations. Evolution proceeds slowly and smoothly over
millions of years
Punctuated Equilibrium
Supported by the work of de Vries
Punctuated Equilibrium
Most species exhibit little netchange in their geological history. Stasis: a state where a species remains unchanged for a long period of time
Evolutionary change happens in a rare and rapid bursts
Fossil Record says…
Gradualism
◦ Transitional forms are
lost due to rarity of formation of fossils
◦ Leads to the observed
gaps in phylogeny
Punctuated Equilibrium
◦ Transitional forms are not
lost; they don’t exist
◦ The jumps observed in