Contributions to the knowledge of Norwegian Tachinidae (Diptera, Oestroidea) Part 1, including an updated checklist

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Contributions to the knowledge of Norwegian Tachinidae (Diptera,

Oestroidea) – Part 1, including an updated checklist

HÅKON HARALDSEIDE

Haraldseide, H. 2012. Contributions to the knowledge of Norwegian Tachinidae (Diptera, Oestroidea) – Part 1, including an updated checklist. Norwegian Journal of Entomology 59, 43–58.

Records are given for 215 specimens representing 34 genera and 41 species of tachinid flies from western Norway. Gymnocheta magna Zimin, 1958, Prooppia nigripalpis (Robineau-Desvoidy, 1847) and Tachina ursina Meigen, 1824 are recorded in Norway for the first time, notes on their identification, biology and distribution are given. An updated checklist of Norwegian Tachinidae with references to sources of records succeeding Rognes (1986) is also given.

Key words: Diptera, Tachinidae, Gymnocheta magna, Prooppia nigripalpis, Tachina ursina, new records, checklist, Norway.

Håkon Haraldseide, Ålavikvegen 4, 4250 Kopervik, Norway. E-mail: hharaldseide@gmail.com

Introduction

Tachinidae is a speciose and morphologically diverse family of flies with members ranging in size from 20 (Tachina grossa (Linnaeus, 1758) (Figure 1)) to only a couple of millimetres. More than 1600 species are known from the Palaearctic Region (Tschorsnig & Richter 1998). Fauna Europaea (Tschorsnig et al. 2004) lists 875 species for Europe and 382 for Scandinavia, 358 and 279 species for Sweden and Finland, respectively. Only 200 species are listed for Norway. In the updated checklist below the total number of species is 220. Undoubtedly many species still await discovery in Norway.

The typical tachinid fly is medium sized, black, greyish dusted and very bristly, usually easily recognized, even in the field. Almost all tachinids share the exclusive character of an enlarged subscutellum.

All tachinids are obligate parasitoids of arthropods, most species attack lepidopteran larvae, but other groups are also parasitized (Coleoptera, Heteroptera, Hymenoptera and Dermaptera to mention the most important).

Tschorsnig & Richter (1998) explains the morphology, biology and behaviour, classification and distribution of Tachinidae in the Palaearctic Region.

Material and methods

In this first in a series of papers on Norwegian Tachinidae I aim to contribute to the knowledge of Tachinidae in Norway by presenting an updated checklist and 215 records from Norway (Figure 2) representing 41 species in 34 genera, including three new species for Norway; Gymnocheta magna Zimin, 1958, Prooppia nigripalpis (Robineau-Desvoidy, 1847) and Tachina ursina Meigen, 1824.

The checklist is based on the previous list by Rognes (1986) and succeeding papers reporting tachinid species new to Norway. Additions since Rognes (1986) are marked with a number which refers to the relevant publication.

Generic and specific synonyms and emendations are only included where the source-records do not agree with the classification

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FIGURE 1. Tachina grossa (Linnaeus, 1758) female. Drawing: Håkon Haraldseide.

FIGURE 2. Collecting sites. Map source: Statens Kartverk

followed here. For earlier names used in Scandinavian literature refer to Rognes (1986). Species are presented alphabetically by subfamily, genus, subgenus (where applicable) and species, tribes are omitted.

Records are presented by: ‘Faunistic Region (Økland 1981), Municipality, Location and European Invertebrate Survey grid-number (Endrestøl 2005).

Remarks on ecology represent general observations as seen in location(s) listed under ‘Material:’. Additional remarks are given when seen useful, including previous records of rare species or species just recently published new to Norway. Extensive notes are only given for species new to Norway.

Identifications follow Tschorsnig & Herting (1994) and Andersen (1996) (except in the case of Siphona geniculata (De Geer, 1776) see Herting et al. (1999)). The identities of several specimens have been confirmed by Theo Zeegers (Soest, The Netherlands).

A somewhat conservative approach to tachinid systematics have been chosen, which, for the

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most part, agrees with current European authors (e.g. Andersen 1996, Tschorsnig et al. 2004). All specimens have been collected by the author and are kept in the author’s private collection unless otherwise noted.

The checklist

Codes: * = new to Norway, 1 = Ødegaard et al. (2009), 2 = Bergström (2008), 3 = Haraldseide (2011), 4 = Ødegaard et al. (2011), 5 = Rognes (2006), 6 = Gammelmo & Sagvolden (2007), 7 = Rognes & Hansen (1996), 8 = Ziegler & Menzel (2000), 9 = Andersen (1996), 10 = Bergström (2007), 11 = Rognes (1995),

Annotations: A1: Christer Bergström (Uppsala, Sweden) has seen material from Norway that belongs to a species that differs from the typical T. nigripes. A2: Gymnosoma rotundatum (Linnaeus, 1758) was published from Norway in Ødegaard et al. (2009) but closer inspection of the genitals has shown that the specimens represent Gymnosoma nudifrons Herting, 1966 (pers. comm. Frode Ødegaard (Trondheim, Norway)).

DEXIINAE

Athrycia Robineau-Desvoidy, 1830 curvinervis (Zetterstedt, 1844) impressa (van der Wulp, 1869) trepida (Meigen, 1824) Billaea Robineau-Desvoidy, 1830 triangulifera (Zetterstedt, 1844)

Blepharomyia Brauer & Bergenstamm, 1889 pagana (Meigen, 1824) piliceps (Zetterstedt, 1859) Campylocheta Rondani, 1859 inepta (Meigen, 1824) praecox (Meigen, 1824) Chaetovoria Villeneuve, 1920 antennata (Villeneuve, 1920) Cyrtophloeba Rondani, 1856 Cyrtophleba, unjust. emendation ruricola (Meigen, 1824) Dexia Meigen, 1826 vacua (Fallén, 1817) Dinera Robineau-Desvoidy, 1830 carinifrons (Fallén, 1817) ferina (Fallén, 1817) grisescens (Fallén, 1817) Dufouria Robineau-Desvoidy, 1830 chalybeata (Meigen, 1824) nigrita (Fallén, 1810)1 Eriothrix Meigen, 1803 prolixa (Meigen, 1824) rufomaculata (De Geer, 1776) Estheria Robineau-Desvoidy, 1830 bohemani (Rondani, 1862) Freraea Robineau-Desvoidy, 1830 gagatea Robineau-Desvoidy, 1830 Klugia Robineau-Desvoidy, 1863 marginata (Meigen, 1824) Periscepsia Gistel, 1848 carbonaria (Panzer, 1798)1 Peteina Meigen, 1838 erinaceus (Fabricius, 1794) Phyllomya Robineau-Desvoidy, 1830 volvulus (Fabricius, 1794)

Prosena Le Peletier & Serville, 1828 siberita (Fabricius, 1775) Ramonda Robineau-Desvoidy, 1863 latifrons (Zetterstedt, 1844) prunaria (Rondani, 1861) ringdahli (Villeneuve, 1922) spathulata (Fallén, 1820) Rondania Robineau-Desvoidy, 1850 dimidiata (Meigen, 1824) fasciata (Macquart, 1834)

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Thelaira Robineau-Desvoidy, 1830 ?nigripes (Fabricius, 1794)A1

solivaga (Harris, 1780)

Trafoia Brauer & Bergenstamm, 1893 monticola Brauer & Bergenstamm, 18931

Trixa Meigen, 1824 Murana Meigen, 1824 (syn.) alpina Meigen, 1824

caerulescens Meigen, 1824 (syn.) conspersa (Harris, 1776)

variegata Meigen, 1824 (syn.)

Voria Robineau-Desvoidy, 1830 ruralis (Fallén, 1810) Wagneria Robineau-Desvoidy, 1830 alpina Villeneuve, 1910 costata (Fallén, 1815) EXORISTINAE

Admontia Brauer & Bergenstamm, 1889 Trichopareia Brauer & Bergenstamm, 1889 (syn.) blanda (Fallén, 1820) grandicornis (Zetterstedt, 1849) Allophorocera Hendel, 1901 ferruginea (Meigen, 1824) Aplomya Robineau-Desvoidy, 1830 confinis (Fallén, 1820) Belida Robineau-Desvoidy, 1863 angelicae (Meigen, 1824) Blondelia Robineau-Desvoidy, 1830 nigripes (Fallén, 1810) Botria Rondani, 1856 Bothria, unjust. emendation frontosa (Meigen, 1824) subalpina Villeneuve, 1910 Brachicheta Rondani, 1861 strigata (Meigen, 1824) Cadurciella Villeneuve, 1927 tritaeniata (Rondani, 1859) Carcelia Robineau-Desvoidy, 1830 atricosta Herting, 1961 lucorum (Meigen, 1824) Chetogena Rondani, 1856 obliquata (Fallén, 1810) Cyzenis Robineau-Desvoidy, 1863 albicans (Fallén, 1810) Drino Robineau-Desvoidy, 1863 galii (Brauer & Bergenstamm, 1891)1

inconspicua (Meigen, 1830)1 lota (Meigen, 1824) vicina (Zetterstedt, 1849) Elodia Robineau-Desvoidy, 1863 ambulatoria (Meigen, 1824) morio (Fallén, 1820) Epicampocera Macquart, 1849 succincta (Meigen, 1824) Erynnia Robineau-Desvoidy, 1830 ocypterata (Fallén, 1810) Eumea Robineau-Desvoidy, 1863 linearicornis (Zetterstedt, 1844) mitis (Meigen, 1824)1 Exorista Meigen, 1803

Subgenus Adenia Robineau-Desvoidy, 1863 mimula (Meigen, 1824)

rustica (Fallén, 1810) Subgenus Exorista Meigen, 1803 fasciata (Fallén, 1820) Frontina Meigen, 1838 laeta (Meigen, 1824) Gastrolepta Rondani, 1862 anthracina (Meigen, 1826)1 Gonia Meigen, 1803 capitata (De Geer, 1776) ornata Meigen, 1826

picea (Robineau-Desvoidy, 1830) sicula Robineau-Desvoidy, 1830 (syn.)

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Hubneria Robineau-Desvoidy, 1847 Huebneria, unjust. emendation

affinis (Fallén, 1810) Istocheta Rondani, 1859 longicornis (Fallén, 1810) Leiophora Robineau-Desvoidy, 1863 innoxia (Meigen, 1824) Lydella Robineau-Desvoidy, 1830 ripae (Brischke, 1885) stabulans (Meigen, 1824) Madremyia Townsend, 1916 clausa (Villeneuve, 1937) Medina Robineau-Desvoidy, 1830 collaris (Fallén, 1820) luctuosa (Meigen, 1824) separata (Meigen, 1824) Meigenia Robineau-Desvoidy, 1830 dorsalis (Meigen, 1824) mutabilis (Fallén, 1810) Myxexoristops Townsend, 1911 arctica (Zetterstedt, 1838)2

blondeli authors, not Robineau-Desvoidy, 1830 Misident- ification (see Bergström 2008)

stolida (Stein, 1924) Nemorilla Rondani, 1856 maculosa (Meigen, 1824) Nilea Robineau-Desvoidy, 1863 hortulana (Meigen, 1824) innoxia Robineau-Desvoidy, 1863 rufiscutellaris (Zetterstedt, 1859) Ocytata Gistel, 1848 pallipes (Fallén, 1820)

Onychogonia Brauer & Bergenstamm, 1889 cervini (Bigot, 1881) flaviceps (Zetterstedt, 1838) Oswaldia Robineau-Desvoidy, 1863 muscaria (Fallén, 1810) spectabilis (Meigen, 1824) Pales Robineau-Desvoidy, 1830 pavida (Meigen, 1824)3 Paracraspedothrix Villeneuve, 1919 montivaga Villeneuve, 19191 Periarchiclops Villeneuve, 1924 scutellaris (Fallén, 1820) Phebellia Robineau-Desvoidy, 1846 glauca (Meigen, 1824) glaucoides Herting, 1961 stulta (Zetterstedt, 1844) villica (Zetterstedt, 1838) Phorocera Robineau-Desvoidy, 1830 obscura (Fallén, 1810) Phryxe Robineau-Desvoidy, 1830 magnicornis (Zetterstedt, 1838) nemea (Meigen, 1824)1 vulgaris (Fallén, 1810) Platymya Robineau-Desvoidy, 1830 fimbriata (Meigen, 1824) Policheta Rondani, 1856 unicolor (Fallén, 1820) Prooppia Townsend, 1926 * nigripalpis (Robineau-Desvoidy, 1847) strigifrons (Zetterstedt, 1838)

Pseudoperichaeta Brauer & Bergenstamm, 1889 palesioidea (Robineau-Desvoidy, 1830) Rhaphiochaeta Brauer & Bergenstamm, 1889

breviseta (Zetterstedt, 1838) Senometopia Macquart, 1834

pollinosa (Mesnil, 1941) separata (Rondani, 1859)

bombycivora Robineau-Desvoidy, 1830 (syn.)

Smidtia Robineau-Desvoidy, 1830 Timavia Robineau-Desvoidy, 1863 (syn.)

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conspersa (Meigen, 1824)4

Staurochaeta Brauer & Bergenstamm, 1889 albocingulata (Fallén, 1820) Tlephusa Robineau-Desvoidy, 1863 cincinna (Rondani, 1859) Winthemia Robineau-Desvoidy, 1830 bohemanni (Zetterstedt, 1844) erythrura (Meigen, 1838) quadripustulata (Fabricius, 1794) Xylotachina Brauer & Bergenstamm, 1891

diluta (Meigen, 1824) Zaira Robineau-Desvoidy, 1830 cinerea (Fallén, 1810) Zenillia Robineau-Desvoidy, 1830 libatrix (Panzer, 1798) PHASIINAE Catharosia Rondani, 1868 pygmaea (Fallén, 1815) Cinochira Zetterstedt, 1845 atra Zetterstedt, 1845 Cistogaster Latreille 1829 globosa (Fabricius, 1775) Cylindromyia Meigen, 1803

Subgenus Cylindromyia Meigen, 1803 brassicaria (Fabricius, 1775) Subgenus Neocyptera Townsend, 1916

interrupta (Meigen, 1824) Subgenus Ocypterula Rondani 1856

pusilla (Meigen, 1824) Gymnosoma Meigen, 1803 clavatum (Rohdendorf, 1947) nudifrons Herting, 1966 Lophosia Meigen, 1824 fasciata Meigen, 1824 Phania Meigen, 1824 curvicauda (Fallén, 1820)4 thoracica Meigen, 1824 Phasia Latreille, 1804

Alophora Robineau-Desvoidy, 1830 (syn.) aurulans Meigen, 18245 hemiptera (Fabricius, 1794)6 obesa (Fabricius, 1798) pusilla Meigen, 1824 Subclytia Pandellé, 1894 rotundiventris (Fallén, 1820) TACHININAE Actia Robineau-Desvoidy, 1830 crassicornis (Meigen, 1824) dubitata Herting, 1971 (syn.) lamia (Meigen, 1838) maksymovi Mesnil, 1952 nigroscutellata Lundbeck, 1927 pilipennis (Fallén, 1810) resinellae (Schrank, 1781) nudibasis Stein 1924 (syn.)

Anthomyiopsis Townsend, 1916 nigrisquamata (Zetterstedt, 1838) Aphantorhaphopsis Townsend, 1926 Asiphona Mesnil, 1954 (syn.)

verralli (Wainwright, 1928) Aphria Robineau-Desvoidy, 1830 longilingua Rondani, 1861 longirostris (Meigen, 1824) Appendicia Stein, 1924 truncata (Zetterstedt, 1838) Ceranthia Robineau-Desvoidy, 1830 abdominalis (Robineau-Desvoidy, 1863)1 lichtwardtiana (Villeneuve, 1931)9 tristella (Herting, 1966)9 verneri (Andersen, 1996)9 Ceromya Robineau-Desvoidy, 1830 bicolor (Meigen, 1824) silacea (Meigen, 1824)7

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Cleonice Robineau-Desvoidy, 1863 callida (Meigen, 1824) nitidiuscula (Zetterstedt, 1859)8 Demoticus Macquart, 1854 plebejus (Fallén, 1810) Dexiosoma Rondani, 1856 caninum (Fabricius, 1781) Eloceria Robineau-Desvoidy, 1863 delecta (Meigen, 1824)1 Entomophaga Lioy, 1864 nigrohalterata (Villeneuve, 1921)9 Eurithia Robineau-Desvoidy, 1844 anthophila (Robineau-Desvoidy, 1830) caesia (Fallén, 1810) connivens (Zetterstedt, 1844) consobrina (Meigen, 1824) intermedia (Zetterstedt, 1844)1 vivida (Zetterstedt, 1838) Germaria Robineau-Desvoidy, 1830 ruficeps (Fallén, 1820) Gymnocheta Robineau-Desvoidy, 1830 * magna Zimin, 1958 viridis (Fallén, 1810)

Hyalurgus Brauer & Bergenstamm, 1893 crucigera (Zetterstedt, 1838)

lucidus (Meigen, 1824) Leskia Robineau-Desvoidy, 1830

aurea (Fallén, 1820)

Linnaemya Robineau-Desvoidy, 1830

Subgenus Linnaemya Robineau-Desvoidy, 1830 vulpina (Fallén, 1810)

Subgenus Ophina Robineau-Desvoidy, 1863 perinealis Pandellé, 1895

Loewia Egger, 1856 erecta Bergström, 200710

phaeoptera authors, not Meigen, 1824. Misidentification (see Bergström 2007) foeda (Meigen, 1824) Lydina Robineau-Desvoidy, 1830 aenea (Meigen, 1824) Lypha Robineau-Desvoidy, 1830 dubia (Fallén, 1810) ruficauda (Zetterstedt, 1838) Macquartia Robineau-Desvoidy, 1830 nudigena Mesnil, 1972 tenebricosa (Meigen, 1824) Nemoraea Robineau-Desvoidy, 1830 pellucida (Meigen, 1824) Nowickia Wachtl, 1894 alpina (Zetterstedt, 1849) marklini (Zetterstedt, 1838) Panzeria Robineau-Desvoidy, 1830 Ernestia Robineau-Desvoidy, 1830 (syn.)

rudis (Fallén, 1810) Pelatachina Meade, 1894 tibialis (Fallén, 1810) Peleteria Robineau-Desvoidy, 1830 rubescens (Robineau-Desvoidy, 1830) Peribaea Robineau-Desvoidy, 1863 hertingi Andersen, 19969 setinervis (Thomson, 1869) fissicornis Strobl, 1909 (syn.)

Phytomyptera Rondani, 1845 Elfia Robineau-Desvoidy, 1850 (syn.)

cingulata (Robineau-Desvoidy, 1830) minutissima (Zetterstedt, 1844) nigrina (Meigen, 1824) zonella (Zetterstedt, 1844)1 Pseudopachystylum Mik, 1891 goniaeoides (Zetterstedt, 1838) Siphona Meigen, 1803 boreata Mesnil, 1960 collini Mesnil, 1960 confusa Mesnil, 1961

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cristata (Fabricius, 1805) flavifrons Stæger, 1849 geniculata (De Geer, 1776) grandistylum Pandellé, 1894 hokkaidensis Mesnil, 1957 nigricans authors, not Villeneuve, 1930 ingerae Andersen, 198211 maculata Stæger, 1849 mesnili Andersen 1982 paludosa Mesnil, 1960 pauciseta Rondani, 1865 rossica Mesnil, 1961 setosa Mesnil, 1960 subarctica Andersen, 19969 variata Andersen, 19829 Solieria Robineau-Desvoidy, 1848 pacifica (Meigen, 1824) Tachina Meigen, 1803

Subgenus Eudoromyia Bezzi, 1906 fera (Linnaeus, 1761)

”Tachina sp.”

magnicornis authors, not Zetterstedt, 1844. Misidentifi- cation (see Novotná et al. 2009)

Subgenus Servillia Robineau-Desvoidy, 1830 *ursina Meigen,

Subgenus Tachina Meigen, 1803 grossa (Linnaeus, 1758) Triarthria Stephens, 1829

setipennis (Fallén, 1810) spinipennis Meigen, 1824 (syn.)

Trichactia Stein, 1924

pictiventris (Zetterstedt, 1855)7

Zophomyia Macquart, 1835 temula (Scopoli, 1763)

Species added to the Norwegian list *Gymnocheta magna Zimin, 1958 (Figures 4B & 5D).

The genus Gymnocheta Robineau-Desvoidy, 1830 is represented in the Oriental, Nearctic and Palaearctic regions (O’Hara 2011), in the latter by eight species (Tschorsnig & Richter 1998). Only

two species are known from Europe; Gymnocheta viridis (Fallén, 1810) and Gymnocheta magna Zimin, 1958, both found in Sweden and Finland but until now only G. viridis has been published for Norway (Rognes 1981, 1986, Tschorsnig et al. 2004).

Like most members of the subfamily Tachininae, Gymnocheta does not lay eggs directly on the host, but in its vicinity and the larvae actively searches for hosts (Tschorsnig & Richter 1998). No hosts for G. magna are known. G. magna is associated with moorland and peatbogs; it flies early May to end of June in Central Europe, later than G. viridis but the flight-period overlaps (Tschorsnig & Herting 1994, Zeegers et al. 2001).

Identification. The most striking feature of the genus Gymnocheta is the shiny metallic colour, a character it shares only with Chrysosomopsis Townsend, 1916 in European tachinids. Usually green but sometimes blue, bronze red or violet. Gymnocheta can be recognized by combining the following features:

Head. Arista bare, cheeks bare, eyes hairy, palps black or dark brown (yellow in Chrysosomopsis).

Thorax. 3+4 dc (3+3 dc in Chrysosom-opsis).

Wing. Vein m angled before wing margin, R5 open, deflection of m with an appendix about as long as r-m (without or only with a tiny appendix in Chrysosomopsis).

G. magna can be separated from G. viridis by combining the following features: Mouth-edge protruding beyond the base of the vibrissae and palps not dilated (Figure 4), femora metallic green in their basal two thirds, abdomen with prone hairs. Males: Parafrontalia in a stripe along the row of the frontal bristles metallic-green, notch of sternite 5 only about as long as its greatest width. Material. RY, Karmøy: Bygnes (EIS 13) 7 July 2011, 1♀; 15 July 2011, 1♀.

Remarks. Specimens were collected on a small stretch of Anthriscus flowers growing on a hilltop bordering a lake to the west and quarry to the east.

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*Prooppia nigripalpis (Robineau-Desvoidy, 1847)

(Figures 3 & 5G).

Phebellia sensu lato (Herting & Dely-Draskovits 1993: 184) was split by Herting (1960: 68) and Wood (1987: 1207) into two genera based on substantial differences in the reproductive mode of the females: species assigned to Phebellia Robineau-Desvoidy, 1846 have ovolarviparous females and species assigned to Prooppia Townsend, 1926 have oviparous females. The external similarities between Phebellia and Prooppia appear to be the result of convergence (Cerretti 2010: 182). This distinction between Phebellia and Prooppia has been adopted by some recent authors for Palaearctic species (e.g., Richter & Wood 1995, Richter 2010, Tachi & Shima 2010, Cerretti 2010) and is followed here.

Prooppia is known from the Palaearctic and Nearctic Regions (O’Hara 2011), there are two species known from Europe; Prooppia nigripalpis (Robineau-Desvoidy, 1847) and

FIGURE 3. Prooppia nigripalpis (Robineau-Desvoidy, 1847) male.

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FIGURE 5A–H. A. Cinochira atra Zetterstedt, 1845 male. B. Phasia hemiptera (Fabricius, 1794) male. C. Trafoia monticola Brauer & Bergenstamm, 1893 male. D. Gymnocheta magna Zimin, 1958 female. E. Siphona ingerae Andersen, 1982 male. F. Pales pavida (Meigen, 1824) male. G. Prooppia nigripalpis (Robineau-Desvoidy, 1847) male. H. Staurochaeta albocingulata (Fallen, 1820) male.

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Prooppia strigifrons (Zetterstedt, 1838), both are known from Sweden and Finland, but until now only Prooppia strigifrons have been recorded from Norway (Tschorsnig et al. 2004).

No hosts for P. nigripalpis are known (Herting 1960, Tschorsnig & Herting 1994).

Identification. Prooppia are blackish-grey medium sized flies. The genus can be identified by combining the following characters:

Head. Arista bare and thickened at most 2/5 of its length, cheeks bare, eyes haired, gena wider than the cheeks at the level of the antennal base, cheeks narrower than the 3rd antennal segment,

palps black.

Thorax. Subapical scutellar bristles reach at least as far behind as do the apicals, lateral scutellar bristles 0.9–1.1x the length of the basal bristles. Pre-alar bristle longer and stronger than the notopleural bristles. Humeral bristles stand in the shape of a triangle with at most 1–2 much weaker additional bristlets.

Abdomen. tergite 2 hollowed to the posterior edge, tergites 3 and 4 with discal bristles.

Wings. Vein m angled before wing margin but no appendix, cell R5 open.

Legs. Middle tibia with 2–3 anterodorsals, hind tibia with 2 dorsal apical spurs.

P. nigripalpis can be separated from P. strigifrons by combining the following: mouthedge not protruding (Figure 3) and tergites dusted ½–¾ of their length.

Material. RY, Karmøy: Slettavatnet (EIS 13) 11 August 2011, 1♂.

Remarks. Sheltered meadow surrounded by tall conifers and deciduous forest, low vegetation. *Tachina ursina Meigen, 1824

The genus Tachina Meigen, 1803 is known form the Oriental, Nearctic, Neotropical and Palaearctic regions (O’Hara 2011). In Europe the genus is represented by 13 species (Novotná et al. 2009). Tachina ursina Meigen, 1824 belongs to the subgenus Servillia Robineau-Desvoidy, 1830 of which there are two additional species known from Europe: Tachina lurida (Fabricius 1781) and Tachina nigrohirta (Stein 1924), only T. ursina is known from Sweden and Finland.

T. ursina is associated with warm and dry

areas, especially forest edges. It flies from end of March to mid-May in in Central Europe.

No hosts for T. ursina are known (Tschorsnig & Herting 1994).

Identification. Tachina species are large flies (9–20 mm) that can be recognized by combining the following:

Head. Arista bare, cheeks hairy, eyes bare, ocellar bristles present, palps thin, few hairs or bristles above the vibrissa, 3rd antennal not longer

than the narrow 2nd.

Thorax. Prosternum bare, propleuron hairy. Abdomen. Often partly red or yellow.

Wings. Vein m angled before wing margin, R5 without a petiole.

Legs. Hind coxae posterodorsally haired. European species of the subgenus Servillia all have a similar habitus to that of a bee; thorax with long hairs as opposed to the typical bristles only. Tachina ursina can be separated from T. lurida and T. nigrohirta by combining the following: Thorax: Dorsal hairs pale yellow or whitish (predomenantly black in T. nigrohirta). Crossed apical scutellar bristles missing or hair-like (Crossed apicals usually strong in T. lurida). 2 or 3 sternopleural bristles (1 (seldom 0 or 2) in T. lurida). Abdomen: Tergite 5 at the anterior edge almost always with a narrow band of dusting (missing in T. lurida). Marginal bristles of tergite 4 as a rule shorter than the segment (longer in T. nigrohirta).

Material. Ø, Halden: Endetjern (EIS 12) 1 May 2012, 1♂ (leg. Magne Pettersen)

Additional records DEXIINAE

Campylocheta praecox (Meigen, 1824)

Material. RY, Karmøy: Skår (EIS 13) 25 March 2011, 1♂; 9 April 2011, 1♀; 25 April 2011, 1♀.

Remarks. Deciduous woodland, tree trunks. Eriothrix rufomaculata (De Geer, 1776)

Material. RY, Karmøy: Grodvatnet (EIS 13) 6 July 2011, 2♂♂; Kopervik 26 August 2011, 1♀;

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Skår: 28 June 2011, 1♀; Slettavatnet: 8 September 2011, 1♀; Visnes: 19 August 2005, 2♀♀ (leg./coll. Magne-Henrik Velde); Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 1♀.

Remarks. Meadows and gardens, flowers and low vegetation.

Trafoia monticola Brauer & Bergenstamm, 1893

(Figure 5C)

Material. RY, Karmøy: Slettavatnet (EIS 13) 10 August 2011, 1♂; 18 August 2011, 2♀♀; 20 August 2011, 1♀ (coll. Theo Zeegers); 23 August 2011, 1♀ (coll. Theo Zeegers).

Remarks. A rare species, just recently recorded in Norway for the first time (Ødegaard et al. 2009). Meadow surrounded by conifers and deciduous forest, bare ground and stones. Voria ruralis (Fallen, 1810)

Material. RY, Karmøy: Bygnes (EIS 13) 8 September 2011, 1♀; Slettavatnet 10 August 2011, 1♂; 17 August 2011, 1♀; 20 August 2011, 2♀♀; 21 August 2010, 1♂2♀♀; 23 August 2011, 2♀♀; 30 August 2011, 3♀♀.

Remarks. Meadows and marshes, low veg-etation.

EXORISTINAE

Admontia blanda (Fallen, 1820)

Material. RY, Karmøy: Skår (EIS 13) 1 August 2011, 1♀.

Remarks. Rural garden.

Allophorocera ferruginea (Meigen, 1824) Material. RY, Karmøy: Bygnes (EIS 13) 7 July 2011, 1♂; 9 July 2011, 1♂; Fotvatnet 25 July 2011, 1♀; Grodvatnet 16 August 2011, 1♂; 5 September 2011, 1♂1♀ (in copula); 24 September 2011, 1♀; Husemyra 23 July 2011, 1♂1♀; Skår 28 June 2011, 1♂; 1 July 2011, 1♂; Slettavatnet 10 August 2011, 1♂; 18 August 2011, 2♀♀; 30 August 2011, 1♀; Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 1♂.

Remarks. Meadows, heathland and gardens, low vegetation and flowers.

Belida angelicae (Meigen, 1824)

Material. RY, Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 2♂♂.

Cyzenis albicans (Fallen, 1810)

Material. HOY, Sveio: Møstre (EIS 22) 15 May 2011, 1♀; RY, Karmøy: Skår (EIS 13) 13 April 2011, 1♀; 14 April 2011, 1♂; 17 May 2011, 1♂; 22 May 2011, 1♀.

Remarks. Gardens, bare ground and low vegetation.

Gonia picea (Robineau-Desvoidy, 1830)

Material. RY, Karmøy: Borgaredalen (EIS 13) 22 April 2011, 1 ♀; Bygnes 24 April 2011, 1♀; Grodvatnet 21 April 2011, 1♂.

Remarks. Heathland and roadsides, bare ground and low vegetation.

Hubneria affinis (Fallen, 1810)

Material. RY, Karmøy: Bygnes (EIS 13) 9 July 2011, 1♀.

Remarks. Meadow, low vegetation. Medina collaris (Fallen, 1820)

Material. RY, Karmøy: Bygnes (EIS 13) 03 August 2011, 2♂♂; Grodvatnet 18 June 2011, 1♂; 28 July 2011, 1♀; 13 August 2011, 1♀; 29 August 2011, 1♀; Husemyra 23 July 2011, 1♂; Kopervik 26 August 2011, 1♀; Skår 4 June 2011, 1♂; Slettavatnet 8 August 2011, 1♂.

Remarks. Meadows and shrubby areas, Anthriscus flowers and foliage.

Meigenia mutabilis (Fallen, 1810)

Material. RY, Karmøy: Skår (EIS 13) 6 June 2011,1♂; 2 August 2011, 1♂; 15 August 2011, 1♂; Slettvatnet 11 August 2011, 1♂.

Remarks. Meadows and marshland, foliage. Oswaldia muscaria (Fallen, 1810)

Material. HOY, Sveio: Møstre (EIS 22) 15 May 2011, 1♂2♀♀; RY, Karmøy: Skår (EIS 13) 4 May 2011, 1♂; 13 May 2011, 1♂; 17 May 2011, 1♂; 19 May 2011, 1♂; 25 May 2011, 1♂; 1 June 2011, 1♀; 8 June 2011, 1♀; 10 June 2011, 1♀; 18 June 2011, 1♀; 28 June 2011, 1♂; Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 1♀.

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Remarks. Meadows and gardens, shrubs and low vegetation.

Pales pavida (Meigen, 1824) (Figure 5F)

Material. RY, Karmøy: Grodvatnet (EIS 13) 16 June 2011, 1♀.

Remarks. Just recently recorded in Norway for the first time (Haraldseide 2011). Roadside (mixed woodland), foliage.

Phebellia glauca (Meigen, 1824)

Material. RY, Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 2♂♂.

Platymya fimbriata (Meigen, 1824)

Material. RY, Karmøy: Grodvatnet (EIS 13) 16 September 2011, 1♀; Husemyra 24 June 2011, 1♂; Slettavatnet 29 July 2011, 1♂; 2 August 2011, 1♂1♀ (in copula); 1♂; 11 August 2011, 1♂; 21 August 2011, 1♂.

Remarks. Coniferous forest-edges and meadows, bare ground and low vegetation. Senometopia pollinosa (Mesnil, 1941)

Material. RY, Karmøy: Husemyra (EIS 13) 23 July 2011, 1♂.

Remarks. Abandoned pasture, grass. Staurochaeta albocingulata (Fallen, 1820) (Figure 5H)

Material. RY, Karmøy: Grodvatnet (EIS 13) 28 July 2011, 1♂.

Remarks. Heathland with scattered Juniperus, foliage.

PHASIINAE

Cinochira atra Zetterstedt, 1845 (Figure 5A)

Material. RY, Karmøy: Skår (EIS 13) 1 July 2011, 1♂.

Remarks. Deciduous woodland, low vege-tation and tree trunks.

Gymnosoma nudifrons Herting, 1966

Material. RY, Karmøy: Fotvatnet (EIS 13) 25 July 2011, 2♀♀; Skår 21 July 2010, 1♀;

Slettavatnet 30 July 2011, 1♂1♀; 11 August 2011, 1♀.

Remarks. Meadows and roadsides, Anthriscus flowers.

Phasia hemiptera (Fabricius, 1794) (Figure 5B)

Material. RY, Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 4♂♂2♀♀.

Remarks. Just recently recorded in Norway for the first time (Gammelmo & Sagvolden 2007). Garden, foraging on Persicaria affinis.

TACHININAE

Actia crassicornis (Meigen, 1824)

Material. RY, Karmøy: Grodvatnet (EIS 13) 11 September 2011, 1♀; Husemyra 2 July 2011, 1♀; 23 July 2011, 2♀♀; Slettavatnet 10 August 2011, 1♀; 11 August 2011, 1♂.

Remarks. Meadows and heathland, low vegetation and Anthriscus flowers.

Actia nigroscutellata Lundbeck, 1927

Material. RY, Karmøy: Skår (EIS 13) 15 August 2011, 1♀.

Remarks. Marsh, Anthriscus flower. Actia pilipennis (Fallen, 1810)

Material. RY, Karmøy: Slettavatnet (EIS 13) 10 August 2011, 1♀.

Remarks. Meadow, Anthriscus flower. Dexiosoma caninum (Fabricius, 1781)

Material. RY, Tysvær: Nedstrand (EIS 14) 1 August 2009, 1♂ (leg./coll. Magne-Henrik Velde); 6 August 2011, 1♂ (leg. Leiv Tommas Haugen). Hyalurgus lucidus (Meigen, 1824)

Material. RY, Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 2♂♂.

Remarks. Roadside, Anthriscus flowers. Linnaemya vulpina (Fallen, 1810)

Material. RY, Karmøy: Grodvatnet (EIS 13) 6 July 2011, 1♂; 16 September 2011, 1♀; Husemyra 23 July 2011, 1♀; 22 July 2010, 1♂; Slettavatnet 29 July 2011, 1♂; 20 August 2011, 1♀.

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Remarks. Heathland and meadows, low vegetation, tree trunks and Anthriscus flowers. Loewia foeda (Meigen, 1824)

Material. RY, Karmøy: Bygnes (EIS 13) 7 July 2011, 1♂; 25 July 2011, 1♂; Grodvatnet 29 August 2011, 1♀; 29 September 2011, 1♀; Slettavatnet 11 August 2011, 1♂.

Remarks. Meadows, Anthriscus flowers and low vegetation.

Lydina aenea (Meigen, 1824)

Material. RY, Karmøy: Bygnes (EIS 13) 25 July 2011, 1♂; Skår 11 August 2011, 1♀; 8 September 2011, 1♀; Slettavatnet 29 July 2011, 1♀.

Remarks. Marshes, meadows and heathland, shrubs and low vegetation.

Lypha dubia (Fallen, 1810)

Material. RY, Karmøy: Husmyra (EIS 13) 19 June 2011, 1♀; 30 June 2011, 1♀; Skår 8 April, 2011, 1♂; 9 April 2011, 2♂♂; 11 April 2011, 1♂; 25 April 2011, 1♂1♀; 12 May 2011, 1♂; 17 May 2011, 1♂; 24 June 2011, 1♀.

Remarks. Woodland and gardens, tree trunks, low vegetation and bare ground.

Eurithia connivens (Zetterstedt, 1844)

Material. RY, Karmøy: Skår (EIS 13) 13 August 2010, 1♀; 6 July 2011, 2♂♂; 13 July 2011, 5♂♂; Slettavatnet July 2011, 1♂; 11 August 2011, 1♀; Suldal: Ropeid, Finnvik (EIS 14) 11 July 2011, 2♀♀.

Remarks. Meadows and gardens, Anthriscus flowers and low vegetation.

Siphona confusa Mesnil, 1961

Material. RY, Karmøy: Skår (EIS 13) 28 April 2011, 1♂1♂ (coll. Theo Zeegers); 4 May 2011, 3♂♂; 5 May 2011, 1♂1♀ (in copula); 13 May 2011, 1♀; 14 May 2011, 1♀; 27 May 2011, 1♀.

Remarks. Garden, bare ground and low vegetation, flowers.

Siphona geniculata (De Geer, 1776)

Material. RY, Karmøy: Grodvatnet (EIS 13)

5 September 2011, 3♀♀; 16 September 2011, 1♀; 29 September 2011, 2♀♀; 30 September 2011, 1♀; Skår 6 June 2011, 1♀; 1 August 2011, 1♀; Slettavatnet 18 June 2011, 1♀; 1 August 2011, 1♂; 8 September 2011, 1♀.

Remarks. Meadows and heathland, low vegetation and flowers.

Siphona ingerae Andersen, 1982 (Figure 5E)

Material. RY, Karmøy: Skår (EIS 13) 10 April 2011, 1♂.

Remarks. A rare species (Rognes 1995, Andersen 1996). Rural garden/deciduous forest-edge, bare ground.

Siphona setosa Mesnil, 1960

Material. RY, Karmøy: Grodvatnet (EIS 13) 29 August 2011, 1♀; 18 September 2011, 1♀.

Remarks. Heathland. Tachina fera (Linnaeus, 1761)

Material. RY, Karmøy: Bygnes (EIS 13) 25 July 2011, 1♂; Grodvatnet 16 June 2011, 1♀; Melstokkevatnet 6 July 2010, 1♂ (leg./coll. Magne-Henrik Velde); Slettavatnet 11 July 2010, 1♂; 22 July 2010, 1♂; 29 July 2011, 3♀♀.

Remarks. Meadows, Anthriscus flowers. Tachina grossa (Linnaeus, 1758)

(Figure 1)

Material. RY, Karmøy: Blikshavn (EIS 13) 23 July 2001, 1♂ (leg./coll. Magne-Henrik Velde); Husemyra 14 July 2011, 1♀; 23 July 2011, 1♂; Slettavatnet 23 July 2010, 2♀♀; Tysvær: Yrkje (EIS 14) 8 August 2005, 1♀ (leg./coll. Magne-Henrik Velde).

Remarks. Meadows, Anthriscus flowers. Triarthria setipennis (Fallen, 1810)

Material. HOY, Sveio: Møstre (EIS 22) 15 May 2011, 1♀; RY, Karmøy: Skår (EIS 13) 11– 19 March 2011, 7♂♂2♀♀ Ex Puparia indoors; 17 May 2011, 1♀.

Remarks. Specimens Ex Puparia were collected during the winter. Pupae were found inside the hollow stems of Anthriscus plants, often together with the dead Dermapteran host.

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Acknowledgements. I am very grateful to Theo Zeegers (Soest, The Netherlands) for his patience with my queries, for his comments on the manuscript and for confirming the identity of many specimens, to Christer Bergström (Uppsala, Sweden), James O’Hara (Ottawa, Canada) and Hans-Peter Tschorsnig (Stuttgart, Germany) for being extra helpful and to the following for their kindness and support: Frode Ødegaard (Trondheim, Norway), Jaakko Pohjoismäki (Friedberg, Germany), Knut Rognes (Stavanger, Norway), Leiv Tommas Haugen (Åkrehamn, Norway), Magne Pettersen (Fredrikstad, Norway), Magne-Henrik Velde (Visnes, Norway), Thomas Pape (Copenhagen, Denmark) and Øivind Gammelmo (Grua, Norway).

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Received: 21 March 2012 Accepted: 31 May 2012

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