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2.7   Bioassays

2.7.1  Transwell  migration  assays  

Migration  assays  were  performed  with  MSCs.  10,000  MSCs  were  seeded  on  to  PET  

membrane   transwell   inserts   (BD   Biocoat)   containing   8µm   pores   in   500µl   assay  

media.  The  inserts  were  placed  into  wells  containing  750µl  of  the  appropriate  media  

and  incubated  for  16h.  Following  incubation,  the  cells  that  remained  inside  the  insert   were  removed  using  a  wet  cotton  bud.  Cells  that  had  migrated  to  the  underside  of   the  transwell  inserts  were  fixed  and  stained  with  the  Reastain  Quickdiff  kit  (Reagena)   according   to   the   manufacturers   instructions.   Membranes   were   then   cut   from   the   transwell  inserts  and  mounted  on  glass  slides  in  immersion  oil  for  imaging  using  a   Leica   DMIL   microscope   with   the   acquisition   software   Leica   Application   Suite.   Cells   were   counted   as   an   average   of   10   independent   fields   at   10X   magnification   (figure   4.1)   or   counted   in   15   fields   at   20X   (general   pattern   shown   in   figure   2.2)   and   summated.  Cells  that  over-­‐lapped  fields  on  the  right  side  were  ignored.  

               

Figure  2.2.  15  fields  of  migrated  cells  that  were  counted  at   20X  following  transwell  migration  assays  

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2.7.2  Cell  proliferation  assays  

2.7.2.1  Detection  of  EdU  incorporation  by  flow  cytometry.  

Treatments  were  exposed  to  10μM  5-­‐ethynyl-­‐2´-­‐deoxyuridine  (EdU)  for  the  desired   time  (10  or  16  h).  Following  exposure,  cells  were  trypsinised,  washed  in  1%  BSA  and   fixed   using   4%   paraformaldehyde   for   10   min   at   room   temperature.   Cells   were   washed   once   with   1%   BSA   and   then   washed   with   Click-­‐iT   saponin-­‐based   permeabilisation  and  wash  reagent.  Afterwards,  fixed  samples  were  incubated  with   500μl   Click-­‐iT   reaction   cocktail   (prepared   according   to   manufacturers   instructions)   and  incubated  in  the  dark  for  30  min  at  RT.  Cells  were  washed  with  Click-­‐iT  saponin-­‐

based  permeabilisation  and  wash  reagent  and  re-­‐suspended  in  200µl  to  be  analysed  

using  a  FACScalibur  (BD  Biosciences)  flow  cytometer  on  a  low  flow  rate.    

2.7.2.2   Detection   of   EdU   incorporation   in   direct   and   transwell   co-­‐

cultures.  

AGS-­‐  or  MKN45-­‐dTomato  cells  were  exposed  to  assay  media  for  48h  prior  to  seeding   co-­‐cultures   to   synchronise   AGS/MKN45   cell-­‐cycles.   To   set-­‐up   co-­‐cultures,   2.4x104/cm2  labeled  cancer  cells  with  or  without  4.8x103/cm2  MSCs,  CAMs  or  cancer  

cells   (to   control   for   cell   numbers)   were   seeded   in   triplicate   into   6cm   dishes   with  

assay  media.  Fourteen  hours  later,  10μM  EdU  was  added  to  each  dish  and  incubated   for   10   hours.   Samples   were   prepared   for   analysis   as   described   in   2.7.2.1.   EdU   incorporation  was  only  assessed  in  cells  expressing  dTomato.  

   

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2.7.2.3  3D  growth  assay  

In   vitro   tumour   cell   growth   was   monitored   using   a   3D   growth   assay   based   on   the   methods   published   by   (Sasser   et   al.,   2007).   This   method   involves   seeding   EGFP-­‐ labeled  cells  into  a  3D  matrix  of  phenol  red  free,  growth  factor  reduced  matrigel  (BD   Biosciences),  whereby  relative  growth  can  be  monitored  over  an  extended  period  of   time.  Fluorescently  tagged  cells  were  produced  using  a  lentivirus  expressing  EGFP  at   a   MOI   of   10.   1x105   GFP-­‐tagged   cells   with   or   without   2x104   MSCs/myofibroblasts  

were  suspended  in  400µl  phenol  red-­‐free  αMEM  1%  GlutaMax  2%  FBS  and  4mg/ml   matrigel  and  placed  (in  triplicate)  into  a  well  of  a  96  well  black-­‐walled  tissue  culture   plate  (Porvair  Sciences)  avoiding  bubble  formation.  Cultures  were  incubated  at  37oC  

5%  CO2  for  5  hours  before  the  addition  of  50µl  phenol  red-­‐free  αMEM  1%  GlutaMax  

2%   FBS.   Immediately   after,   a   day   0   fluorescent   reading   using   a   GloMax®-­‐Multi   Detection   System   (Promega)   was   taken.   Relative   fluorescence   units   (RFU)   were   normalised  to  identical  unlabeled  cell  controls  and  the  plate  was  analysed  each  day   for   6   days.   Before   reading,   plates   needed   to   be   cooled   at   RT   for   15   minutes   to   prevent  condensation.    

 

2.7.3  Collagen  contraction  assay  

Cells   were   trypsinised   and   re-­‐suspended   in   DMEM   8%   FBS   at   a   density   of   80,000  

cells/ml.  A  collagen  matrix  solution  was  prepared  on  ice  consisting  of  800µl  collagen  

I  (3.75mg/ml),  100µl  10X  DMEM,  80µl  H2O  and  20µl  1M  NaOH  per  sample.  1ml  cell  

suspension  was  added  to  the  collagen  solution,  mixed  thoroughly  (with  care  taken  to  

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matrices  were  incubated  for  2  hours  to  allow  polymerisation  before  being  detached  

using  a  200µl  tip  and  500µl  serum-­‐free  DMEM  was  added  to  each  well  to  reach  a  

final  FBS  concentration  of  2%.  24hs  later,  wells  and  collagen  matrices  were  imaged   using  an  8  megapixel  digital  camera  and  processed  using  ImageJ.  

 

2.7.4  Adipogenic  induction  

70-­‐90%   confluent   cells   were   cultured   in   adipogenic   induction   medium   (αMEM   supplemented  with  10%  FBS,  1%  P-­‐S,  1%  GlutaMax,  0.1nM  dexamethasone,  0.45mM   IBMX,   10ug   insulin   and   125uM   indomethacin)   for   28   days.   Control   cells   were  

cultured   in   αMEM   supplemented   with   10%   FBS,   1%   P-­‐S,   1%   GlutaMax.   After   adipogenic  induction,  cells  were  fixed  in  4%  formaldehyde  for  1h  at  4oC  followed  by  

gentle   washing   with   MQ   water.   Fixed   cells   were   then   incubated   in   2mls   60%   isopropanol  for  5  minutes  followed  by  addition  of  3  parts  oil  red  O  stock  (3g/L  oil  red   O  in  isopropanol,  sterile  filtered)  and  2  parts  MQ  water  for  a  further  5  mins.  Cells   were  then  carefully  rinsed  in  MQ  water  until  clear  and  imaged  (in  HBSS)  using  a  Leica   DMIL   microscope   with   the   acquisition   software   Leica   Application   Suite   and   processed  using  Adobe  Photoshop  (Fink  and  Zachar,  2011).  

 

2.7.5  Osteogenic  induction  

70-­‐90%   confluent   cells   were   cultured   in   osteo-­‐base   medium   (OBM;   αMEM  

supplemented   with   20%   FBS,   1%   PS,   2%   GlutaMax,   5mM  β-­‐glycerophosphate   and  

50µg/ml   ascorbate-­‐2-­‐phosphate)   for   7   days   followed   by   culture   in   osteo-­‐

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21  days  (control  cells  were  cultured  in  αMEM  supplemented  with  20%  FBS,  1%  P-­‐S,   2%   GlutaMax).   Cells   were   fixed   for   1h   at   4oC   in   4%   formaldehyde   and   calcium  

deposits  were  stained  with  40mM  alizarin  red  (pH  4.1).  Images  were  taken  on  a  Leica   DMIL   microscope   using   the   acquisition   software   Leica   Application   Suite   and   processed  using  Adobe  Photoshop  (Krause  et  al.,  2011).