The maculatus group

Diagnosis. Elytron in some species with round yellowish spot. Apex of female protibia without premucro. TA in lateral aspect similar as in fig. 13, without distinct basal sclerite.

Euops maculatus VOSS, 1924

Euops maculata Voss, 1924: 48. Complete bibliography in appendix 1.

(Figs. 40, 96-97, 124-125, 140, 163, 172, 186, 203, 215, 230, 244)

Diagnosis. Coloration of elytron bluish, in anterior half with oblique yellowish spot [rarely missing] (Fig. 40). Pronotum microreticulate, anteriorly at middle distinctly swollen. Male: TA as in fig. 163. Female: Apex of protibia without premucro. Setose patch of venter consisting of three double rows of modified setae. Ovipositor without styli.

Range. Fig. 244. Sorong Prov. (Mysol Isl., Salawatti Isl., Makbon), Manokwari Prov. (Wandammen Bay), Jayawijaya Prov. (Samboka), Western Prov. (Kiunga), Milne Bay Prov. (Normanby Isl.). Altitude: 30-200 m.

Type material. Lectotype, male (by present designation): IRIAN JAYA, Sorong Prov., Mysol; labels: 1)

[white, rectangular] "Mysol, Wallace [handwritten]" 2) [white with black bordering, rectangular] "fulvescens Blik.

[handwritten]" 3) [white, rectangular] "Coll. Haag [printed]" 4) [white with black bordering, rectangular] "Euops

maculata n. sp. [handwritten], det. E. Voß [printed]" 5) [white, rectangular] "coll. DEI Eberswalde [printed]" 6) [red,

rectangular] "Syntypus [printed]" 7) [red, rectangular] "Lectotype, Euops maculatus Voss, det. A.Riedel 2001

[printed]" (DEI). Paralectotypes: IRIAN JAYA, Sorong Prov., Mysol: 1 female; labels: 1) [white, rectangular] "Mysol, Wallace [handwritten]" 2) [white, rectangular] "Coll. Haag [printed]" 3) [red, rectangular] "Syntypus

[printed]" 4) [white, rectangular] "Euops maculata m. [handwritten] 5) [white, rectangular] "coll. DEI Eberswalde

[printed]" 6) [red, rectangular] "Paralectotype, Euops maculatus Voss, det. A.Riedel 2001 [printed]" (DEI).

Other material examined: IRIAN JAYA, Sorong Prov.: 27 males, 21 females, Salawatti Isl., Kalobo, ca. 10-30 m, 19-22.X.1996, leg. A.Riedel (ARC); 1 female, Salawatti Isl., Kalyam, 0-380 m, 3.XI.1996, leg. A.Riedel (ARC); 1 male, 2 females, Salawatti Isl., Waijan, SP2, ca. 50-100 m, 23-25.X.1996, leg. A.Riedel (ARC); 2 males, 1 female, Makbon, Malawor, 50-280 m, 28.I.2001, leg. A.Riedel (ARC); Manokwari Prov.: 4 males, 1 female, Wandammen Bay, Wasior, DMP logging camp, Km 38, Sararti, S 03° 05´ E 134° 27´, 100-200 m, 7-9.I.2001, leg. A.Riedel (ARC); Jayawijaya Prov.: 6 males, 8 females, Samboka, upper Kolff River, ca. 200 m, 10-14.X.1996, leg.

A.Riedel (ARC). PAPUA NEW GUINEA, Western Prov.: 1 male, 5 females, Kiunga, Fly River, 7-8.IX.1957, leg. W.W.Brandt (BPBM); 1 male, Kiunga, Fly River, 1-7.X.1957, leg. W.W.Brandt (BPBM); 1 female, Kiunga, Fly River, 15.VIII.1957, leg. W.W.Brandt (BPBM); 1 female, Kiunga, Fly River, 4-8.VII.1957, leg. W.W.Brandt (BPBM); 1 female, Kiunga, Fly River, 8-10.VIII.1957, leg. W.W.Brandt (BPBM); 1 female, Kiunga, Fly River, 9-14.X.1957, leg. W.W.Brandt (BPBM); 1 female, Kiunga, Fly River, 26-28.X.1957, leg. W.W.Brandt (BPBM); Milne Bay Prov.: 2 males, Normanby Isl., Wakaiuna, Sewa Bay, 1-5.XI.1956, leg. W.W.Brandt (BPBM); 1 male, Normanby Isl., Wakaiuna, Sewa Bay, 1-10.XI.1956, leg. W.W.Brandt (BPBM); 1 female, Normanby Isl., Wakaiuna, Sewa Bay, 21-30.XI.1956, leg. W.W.Brandt (BPBM); 1 male, Normanby Isl., Wakaiuna, Sewa Bay, 1-8.I.1957, leg. W.W.Brandt (BPBM).

Redescription.

Lectotype, male. Length, pronotum + elytron: 2.22 mm. Coloration deep ferruginous to black; elytron with bluish lustre, in anterior half with oblique yellowish creamy colored spot from interval 3 to interval 7; scutellum, elytral humerus, sides of metathorax and anterior surface of procoxa metallic green; antenna yellowish. Head. As in fig. 124. Gena 0.85 X as long as width of head immediately behind eyes. Vertex weakly microreticulate; laterally with shallow, transverse wrinkles. Eyes in dorsal view moderately prominent from lateral contour of head, medially contiguous for ca. 0.5 X their length. Profile of vertex immediately behind eye straight, towards base rounded. Ventral surface of head in front of gular region shining, weakly transversely rugulose, with deep median furrow. Rostrum 2.41 X as long as mouthparts; at widest point 1.46 X wider than at base; in cross section dorsally moderately rounded. Clypeus with median notch. Interantennal area with almost straight lateral margins converging. Profile of rostrum with dorsal contour at interantennal area forming blunt angle, rounded; anteriorly in straight line to apex; ventral contour weakly sinuate, at base weakly concave, towards apex weakly convex; dorsal and ventral contour of rostrum basally subparallel, in apical third converging. Venter of rostrum poorly delimited against venter of head, basally with indistinct submental median carina and with pair of shallow lateral impressions. Mouthparts. Prementum at base 1.6 X wider than long, ca. 3.0 X wider than at apex; with straight lateral margins converging apicad; with three moderately long, acute apical processes, subequal in length. Antenna as in fig. 140. Proventriculus (as in figs. 94- 95) with eight primary gnathal ridges; cutting edge of gnathal ridge in profile of subtriangular shape, rectangular; posteriad of each gnathal ridge with moderately large, denticulate pulvillus. Prothorax 0.90 X as long as wide; with deep subbasal constriction; sides rounded to apex; with shallow preapical constriction; disc dull to weakly shining, microreticulate, with sparse shallow punctures; behind preapical constriction at middle distinctly swollen; in lateral aspect with center of side densely deeply punctate. Metanotum without sutural spines, but with indistinct pair of bluntly angular projections; with pair of lateral lobes. Elytron at humerus simple; 2.23 X longer than wide; striae moderately impressed. Thoracic venter. Prepectus short, subequal to postpectus. Height of pterothorax 0.82 X length of elytron. Procoxa in anterior aspect 1.06 X as long as wide; with weak rectangular extension ventrolateral of articulation with trochanter. Legs. Profemur moderately swollen; strongly asymmetrically

clavate; dorsal and ventral contours straight in basal third, continuing in convex line to apex; anterior contour subangulate, in weakly sinuate line to angulation at apical 0.28, continuing in straight line to apex; posterior contour weakly convex; ventral surface smooth, without protrusion. Protibia evenly curved ventrad; ventral surface crenulate, in basal half subglabrous, in apical half moderately densely setose with short setae; uncus in dorsal position, tibial apex ventrad of uncus obliquely truncate; anterior distal comb complete; posterior distal comb oblique, behind tarsal articulation, dorsally and ventrally shortened. Mesotibia subapically simple, without extension. Abdomen. Pygidium 0.89 X as long as wide; microreticulate; densely, moderately deeply punctate; sparsely setose with inconspicuous, long, recumbent setae. Venter with sparse suberect setae. Genitalia. Sternite VIII as in fig. 203; apex weakly bilobed, with moderately long setae; base concave. Tegminal plate (Fig. 172) with sides subparallel towards apex; apical edge broadly rounded, with dense long setae. Aedeagus. Apical scoop of pedon (Fig. 186) ca. 0.46 X as long as wide, evenly rounded. Tectum with well-sclerotized lateral frame converging with straight sides to truncate apex, medially hyaline. TA as in Fig. 163. Endophallus in apical portion without sclerite.

Paralectotype, female. Same as described male except: length, pronotum + elytron: 2.67 mm. Coloration of body lighter ferruginous. Head. As in fig. 125. Gena 0.67 X as long as width of head immediately behind eyes. Profile of vertex immediately behind eye weakly concave, towards base evenly convex. Rostrum 2.00 X as long as mouthparts; at widest point 1.39 X wider than at base. Profile of rostrum more slender, with dorsal and ventral contour basally subparallel, in apical half converging. Prothorax 0.87 X as long as wide. Elytron 2.15 X longer than wide. Height of pterothorax 0.86 X length of elytron. Procoxa in anterior aspect 0.98 X as long as wide. Legs. Profemur similar as in male, but shorter and stouter; ventral contour convex from base to apex. Protibia with dorsal contour evenly weakly convex; ventral contour sinuate, basally convex, apically concave; ventral surface crenulate, sparsely setose with subrecumbent setae; tibial apex with uncus in dorsal position; without premucro; anterior distal comb complete, ventrally passing base of uncus. Abdomen. Pygidium 0.92 X as long as wide. Setose patch of venter 1.11 X as long as wide, consisting of three double rows of modified setae on sternite III-V with each sternite bearing one double row. Genitalia. Ovipositor without styli. Sternite VIII as in fig. 215; apex subtruncate, laterally rounded; without apodeme. Spermatheca as in fig. 230, compact; with body weakly tapering apicad.

Intraspecific variation. Four geographical forms of uncertain taxonomic status can be distinguished, mainly based on different color patterns. It is possible that some of them belong to distinct, separate species. More material should be available for a comprehensive study, especially from Papua New Guinea.

Length, pronotum + elytron: 2.09-3.25 mm (n=67, =2.64 ± 0.25). Coloration. Shape, position and tint of pale elytral spot exhibiting considerable variation. Typical color form (from Mysol, Salawatti, Sorong, Wasior) with ovate to subrotund yellowish creamy colored spot, usually wider than long, rarely as wide as long, keeping distance to both elytral suture and to humerus. Specimens from Samboka rarely as in typical color form; more often spot smaller (on interval 3 to interval 6) and of darker ferruginous tint, or entirely absent. Elytral spot of Kiunga-specimens usually large, obliquely ovate, longer than wide, coming close to humerus and sometimes also to sutural interval. Elytral spot of Normanby-specimens of ferruginous tint limited to intervals 5 to 6, about twice as long as wide. Primary color of elytron in Samboka-form with blue lustre, as in typical form; in Kiunga-form bright metallic green; in Normanby-form dark ferruginous with violet lustre. Prothorax 0.85-0.96 X as long as wide (n=66, =0.89 ± 0.03); disc in typical form and in Samboka-form with subapical median swelling more distinct in larger specimens, rarely without distinct swelling in very small specimens; Kiunga- and Normanby-form always without such subapical swelling, even in larger specimens. Elytron 2.02-2.29 X longer than wide (n=66, =2.16 ± 0.05). Thoracic venter. Height of pterothorax 0.81-0.89 X length of elytron in males (n=31, =0.85 ± 0.02), 0.83-0.91 X length of elytron in females (n=35, =0.87 ± 0.02). Legs. Profemur of larger males more strongly swollen. Abdomen. Pygidium 0.98-1.11 X wider than long in males (n=21, =1.04 ± 0.05), 1.02-1.23 X wider than long in females (n=28, =1.10 ± 0.05). Female patch of modified setae 0.84-1.26 X as long as wide (n=32, =1.05 ± 0.10).

Euops, new species 16

Range. Jayapura Prov. (Lereh), East Sepik Prov. (Amok). Altitude: 165-300 m.

Material examined: IRIAN JAYA, Jayapura Prov.: 2 males, Lereh, 300-500 m, 26.I.1996, leg. A.Riedel (ARC). PAPUA NEW GUINEA, East Sepik Prov.: 1 male, Amok (N Maprik), 165 m, 6.I.1960, leg. T.C.Maa (BPBM).

Euops, new species 17

Range. Manokwari Prov. (Wandammen Bay). Altitude: ca. 580-950 m.

Material examined: IRIAN JAYA, Manokwari Prov.: 2 males, 1 female, Wandammen Bay, Wondiwoi Mts., Wasior, 580-950 m, 3.I.2001, leg. A.Riedel (ARC).

Euops, new species 18

Range. Paniai Prov. (Epomani). Altitude: ca. 1350-1400 m.

Material examined: IRIAN JAYA, Paniai Prov.: 16 males, 16 female, Epomani, Ugida, Km 179, 1350- 1400 m, 19-20.I.1996, leg. A.Riedel (ARC).

Euops, new species 19

Range. Manokwari Prov. (Wandammen Bay), Paniai Prov. (Nabire). Altitude: 700- 750 m.

Material examined: IRIAN JAYA, Manokwari Prov.: 2 males, 2 females, Wandammen Bay, Wondiwoi Mts., Wasior, 300-700 m, 14.I.2001, leg. A.Riedel (ARC); 1 male, Nabire, Topo, Pusppensaat, Km 54, 750 m, 20- 21.XII.1995, leg. A.Riedel (ARC).

Euops, new species 20

Range. Sorong Prov. (Batanta Isl.), Paniai Prov. (Epomani), Japen-Waropen Prov. (Serui), Jayapura (Cyclops Mts.), Morobe Prov. (Wau), East New Britain Prov. (Keravat). Altitude: 135-1200 m.

Material examined: IRIAN JAYA, Sorong Prov.: 2 males, Batanta Isl., Waylebet, 100-250 m, 31.X.1996, leg. A.Riedel (ARC); Japen-Waropen Prov.: 1 male, 3 females, Japen Isl., Serui, Kontiunai, road to Ambaidiru, 600-700 m, 23.XII.2000, leg. A.Riedel (ARC); 2 females, Japen Isl., Serui, Kontiunai, road to Ambaidiru, 600- 700 m, 25.XII.2000, leg. A.Riedel (ARC); Paniai Prov.: 2 males, Epomani, Km 145, 550-750 m, 15-16.I.1996, leg. A.Riedel (ARC); 1 male, Epomani, Km 169, 900-1100 m, 18.I.1996, leg. A.Riedel (ARC); Jayapura Prov.: 1 male, Cyclops Mts., Ifar, 300-500 m, 29.VI.1962, leg. J.L.Gressitt (BPBM). PAPUA NEW GUINEA, Morobe Prov.: 1 male, Wau, 1200-1300 m, 1-9.IX.1965, leg. J.Sedlacek (BPBM); East New Britain Prov.: 2 males, 1 female, Keravat, 135 m, 20-25.XI.1959, leg. T.C.Maa (BPBM).

Euops, new species 21

Range. Southern Highlands Prov. (L. Kutubu, Karimui), Morobe Prov. (Wau, Sattelberg), Central Prov. (Pt. Moresby). Altitude: 500-1200 m.

Material examined: PAPUA NEW GUINEA, Southern Highlands Prov.: 1 male, Karimui, 1080 m, 13.VII.1963, leg. J.Sedlacek (BPBM); Morobe Prov.: 1 male, Finschhafen, Sattelberg, 100-800 m, 18.X.1992, leg. A.Riedel (ARC); 1 male, Wau, McAdam Park, 1200-1300 m, 7.V.1965, leg. J.Sedlacek (BPBM); 1 male, Wau, 1050-1100 m, 15.XII.1961, leg. J., J.H. & M.Sedlacek & G.Monteith (BPBM); 1 male, Wau, Mt. Missim, 950- 1300 m, 8-9.I.1966, leg. J.&M.Sedlacek (BPBM); Central Prov.: 1 male, Bisianumu, E of Pt. Moresby, 500 m, 8.VI.1955, leg. J.L.Gressitt (BPBM).

Euops, new species 22

Range. Central Prov. (Daradae Plain). Altitude: 500 m.

Material examined: PAPUA NEW GUINEA, Central Prov.: 2 males, Daradae Pl´n, 80 km N to Pt. Moresby, 500 m, sweeping, 6.IX.1959, leg. T.C.Maa (ARC).