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Use of Interstocks and Interstems Interstock or interstem trees are comprised

In document Apples - Botany, Production and Uses (Page 132-139)

Cheryl R. Hampson 1 and Henk Kemp 2

5.11 Use of Interstocks and Interstems Interstock or interstem trees are comprised

of three genetically different components (Fig. 5.6).

Traditionally, interstocks (interstems) were used in raising fruit trees only when

graft compatibility between the rootstock and scion was a problem. This is a common practice when raising certain cultivars of pear on quince rootstocks. Interstock use in apple is less widespread and is generally only applicable when either the desired dwarfing rootstock is difficult to propagate on its own roots (e.g. Ottawa 3) or the soils are unsuited to planting dwarfing rootstocks (e.g. infested with woolly apple aphid or subject to drought or waterlogging). Trunk builders or ‘staddles’ of winter-hardy culti-vars, such as ‘Hibernal’, have been used for a very long time in central Europe to prevent freeze damage to the tree trunk, and cold-tolerance benefits have also been recorded when using interstems of more usual length (Wertheim, 1985).

It is fortunate that, for reasons not understood, a significant part of the dwarf-ing influence of an apple rootstock is attrib-utable to factors associated with its shank (stem piece) rather than its root system.

This means that, by inserting a short length of the dwarfing rootstock as an interstock between a more invigorating rootstock and the scion, the grower can achieve semi-dwarfed trees. Up to approximately 35 cm in length, the longer the interstock stem piece, the more dwarfing the effect (Parry and Rogers, 1972). Trials in Poland have shown increased productivity when using P.2 as an interstock (Kruczynska and Czynczyk, 1998).

Trees raised with interstocks are slightly more expensive to produce but often their

Graft union

Scion

Interstock or interstem

Rootstock

Fig. 5.6. Dwarfing interstock/interstem in relation to apple scion and rootstock.

benefits warrant this extra expenditure by the grower. Further information on raising trees with interstocks (interstems) can be found in Chapter 6.

Occasionally, stem pieces of other scion cultivars (such as ‘Golden Delicious’ or

‘Summerred’) are inserted between the root-stock and the chosen scion. This can improve

the precocity of cropping and winter-hardiness of the tree. Interstocks can also help regulate the vigour of shoot growth on the scions (Wertheim and Callesen, 2001).

Also, in The Netherlands an interstem of

‘Dubbele Zoete Aagt’ is used with the scion cultivar ‘Cox’s Orange Pippin’ to prevent trunk rot caused by P. cactorum.

Further General Reading on Apple Rootstocks

Ferree, D.C. and Carlson, R.F. (1987) Apple rootstocks. In: Rom, R.C. and Carlson, R.F. (eds) Rootstocks for Fruit Crops. Wiley Interscience, New York, pp. 107–144.

Hatton, R.G. (1917) Paradise apple rootstocks. Journal of the Royal Horticultural Society 42, 361–399.

Lockhard, R.G. and Schneider, G.W. (1981) Stock and scion growth relationships and the dwarfing mech-anisms of apple. Horticultural Reviews 3, 315–375.

Sadowski, A. and Hrotko, K. (1999) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, 131 pp.

Tubbs, F.R. (1972) Research fields in the interaction of rootstocks and scions in woody perennials, Parts 1 and 2. HortAbstracts 43, 247–253, 325–335.

Webster, A.D. (1995) Temperate fruit tree rootstock propagation. New Zealand Journal of Crop and Horticultural Science 23, 355–372.

Wertheim, S.J. (1998) Apple rootstocks. In: Rootstock Guide: Apple, Pear, Cherry, European Plum. Fruit Research Station, Wilhelminadorp, The Netherlands, pp. 19–59.

References

Atkinson, C.J., Policarpo, M., Webster, A.D. and Kuden, A.M. (1999) Drought tolerance of apple root-stocks: production and partitioning of dry matter. Plant and Soil 206, 223–235.

Autio, W.R. (1991) Rootstocks affect ripening and other qualities of ‘Delicious’ apples. Journal of the American Society for Horticultural Science 116, 378–382.

Barden, J.A. and Marini, M.E. (1992) Maturity and quality of ‘Delicious’ apples as influenced by root-stock. Journal of the American Society for Horticultural Science 117, 547–550.

Barritt, B.H., Konishi, B.S. and Dilley, M.A. (1995) Performance of three apple cultivars with 23 dwarfing rootstocks during 8 seasons in Washington. Fruit Varieties Journal 49, 158–170.

Bessho, H. and Soejima, J. (1992) Apple rootstock breeding for disease resistance. Compact Fruit Tree 25, 65–72.

Brase, K.D. (1954) Similarity of the Clark dwarf and East Malling rootstock VIII. Proceedings of the American Society for Horticultural Science 61, 95–98.

Callesen, O. (1997) Testing 20 apple rootstocks. Acta Horticulturae 451, 137–145.

Campbell, A.I. and Wilson, D. (1962) Apomictic seedling rootstocks for apples: Progress Report III. In:

Annual Report of Long Ashton Research Station for 1961. Long Ashton, Bristol, UK, pp. 68–70.

Carlson, R.F. (1980) The Michigan apple rootstock clones – an update. Acta Horticulturae 114, 159–169.

Carlson, R.F. and Perry, R.L. (1986) Mark apple rootstock. HortScience 21, 165.

Cummins, J.N. and Aldwinckle, H.S. (1995) Breeding rootstocks for tree fruit crops. New Zealand Journal of Crop and Horticultural Science 23, 395–402.

Czynczyk, A. (1974) The effect of various rootstocks on the field performance and winterhardiness of three apple cultivars. Fruit Science Reports 1, 26–33.

Czynczyk, A. and Olszewska, B. (1990) Growth and yielding of 3 apple cultivars on rootstocks of Polish and foreign breeds. Fruit Science Reports 17, 65–75.

Czynczyk, A. and Piskor, E. (2000) Effect of P.22 rootstocks propagated in vitro and traditionally on growth and fruiting of two apple cultivars. Folia Horticulturae 12(1), 29–39.

Denissen, J.M., Kunneman, B.P.A.M. and Versteegen, F.H.M.M. (1993) Onderstammen uit weefselkweek zijn snel te vermeerderen. Fruitteelt 83(21), 18–19.

Dvorák, A. (1983) Breeding and evaluation of new vegetatively propagated apple rootstocks of J-TE group: progressive trends in fruit production. In: Symposium Hradic Králové 1981, pp. 117–133.

Dvorák, A. (1988) Breeding of rootstocks ‘J-TE’ and their influence on growth and productivity of differ-ent cultivars. Acta Horticulturae 224, 325–329.

Elfving, D.C., Schecter, I. and Hutchinson, A. (1993) The history of the Vineland (V) apple rootstocks.

Fruit Varieties Journal 47, 52–58.

Faby, R., Clever, M. and Tiemann, H. (1986) Freigabe der Apfelunterlagen ‘J.9’. Mitteilungen des Obstbauversuchsringesdes Alten Landes 41, 392–399.

Fernandez, R.T., Perry, R.L. and Flore, J.A. (1997) Drought response of young apple trees on three root-stocks: I. Growth and development. Journal of the American Society for Horticultural Science 122, 14–19.

Ferree, D.C. and Carlson, R.F. (1987) Apple rootstocks. In: Rom, R.C. and Carlson, R.F. (eds) Rootstocks for Fruit Crops. Wiley Interscience, New York, pp. 107–144.

Ferree, D.C., Ellis, M.A. and Hall, F.R. (1983) Tree loss due to fire blight infection of rootstocks and inter-stems in Ohio apple orchards. Compact Fruit Tree 16, 116–120.

Ferree, D.C., Hirst, P.M., Schmid, J.C. and Dotson, P.E. (1995) Performance of three apple cultivars with 22 dwarfing rootstocks during 8 seasons in Ohio. Fruit Varieties Journal 49, 171–178.

Fischer, M. (1997) The Pillnitz apple rootstock breeding and selection results. Acta Horticulturae 451, 89–94.

Fischer, M. (1999) New dwarfing and semi-dwarfing Pillnitz apple rootstocks; Supporter 1, 2, 3 and 4. In:

Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 27–28.

Granger, R.L. (1984) The response of two apple cultivars to different combinations of dwarfing root-stocks, tree densities and training systems. Acta Horticulturae 146, 215–222.

Hatton, R.G. (1917) Paradise apple rootstocks. Journal of the Royal Horticultural Society 42, 361–399.

Hatton, R.G. (1935) Apple rootstock studies: effect of layered stocks upon the vigour and cropping of cer-tain scions. Journal of Pomology and Horticultural Science 13, 293–352.

Hewetson, F.N. (1944) Growth and yield of McIntosh apple trees influenced by the use of intermediate stem pieces. Proceedings of the American Society for Horticultural Science 45, 181–186.

Higgs, K.H. and Jones, H.G. (1991) Water relations and cropping of apple cultivars on a dwarfing root-stock in response to imposed drought. Journal of Horticultural Science 66, 367–379.

Hirst, P.M. and Ferree, D.C. (1995) Effect of rootstock and cultivar on the growth and precocity of young apple trees. Fruit Varieties Journal 49, 34–41.

Hulko, I.P. and Hulko, B.I. (1999) Results of study of clonal apple rootstocks in stoolbeds and in nursery.

In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 41–42.

Jackson, J.E. (1986) M.20 – a super dwarf rootstock for apple. Acta Horticulturae 160, 47–48.

Jakubowski, T. (1999a) Achievements in apple rootstock breeding in Poland. In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp.

47–48.

Jakubowski, T. (1999b) Preliminary evaluation of new apple rootstock clones. Proceedings of the Eucarpia Symposium on Fruit Breeding. Acta Horticulturae 484, 97–100.

Johnson, W.C. (1999) The USDA-ARS/Cornell University apple rootstock breeding and evaluation pro-gram. Compact Fruit Tree 32, 73–76.

Jones, O.P. (1971) Effects of rootstocks and interstocks on the xylem sap concentration in apple trees:

effects on nitrogen, phosphorus and potassium content. Annals of Botany 35, 825–836.

Jones, O.P. (1975) Effect of dwarfing interstocks on the xylem sap composition of apple trees: effect on nitrogen, potassium, phosphorus, calcium and magnesium content. Annals of Botany 40, 1231–1235.

Kamboj, J.S., Blake, P.S., Quinlan, J.D., Webster, A.D. and Baker, D.A. (1997) Recent advances in studies on the dwarfing mechanism of apple rootstocks. Acta Horticulturae 451, 75–82.

Kapichnikova, N.G. (1999) Effect of clonal rootstocks on apple tree performance during first five years after planting. In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards.

Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 49–50.

Kruczynska, D. and Czynczyk, A. (1998) [The effect of dwarf rootstocks and interstems on growth yield and mineral leaf content of three apple cultivars: growth and yield.] Zeszyty Naukowe Instytutu Sadownictwa I Kwiaciarstwa 5, 25–36.

Kuldoshin, I.A. (1999) Perspective clonal rootstocks for intensive orchards in the central areas of Russia.

In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 63–64.

Kurlus, R. and Ugolik, M. (1999) Effect of 13 rootstocks on growth and yielding of Sampion apple trees.

In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 65–66.

Lemoine, J. and Gaudin, J. (1991) Porte-greffe du pommier et sensibilité au Phytophthora cactorum.

L’Arboriculture Fruitière 443, 19–23.

Lockhard, R.G. and Schneider, G.W. (1981) Stock and scion growth relationships and the dwarfing mech-anisms of apple. Horticultural Reviews 3, 315–375.

Lord, W.J., Greene, D.W., Damon, R.A., Jr and Baker, J.H. (1985) Effects of stempiece and rootstock combi-nations on growth, leaf mineral concentrations, yield and fruit quality of Empire apple trees. Journal of the American Society for Horticultural Science 110, 422–425.

Luckwill, L.C. and Campbell, A.I. (1954) The use of apomictic seedlings rootstocks for apples: progress report. In: Annual Report of Long Ashton Research Station for 1953. Long Ashton, Bristol, UK, pp. 47–53.

Mantinger, H. (1996) Eine neue Apfel-Unterlagenserie aus der Tshechei. Obstbau Weinbau 33, 255–256.

Olien, W.C. and Lakso, A.N. (1984) A comparison of the dwarfing character and water relations of five apple rootstocks. Acta Horticulturae 146, 151–157.

Olien, W.C., Ferree, D.C. and Bishop, B.L. (1991) Long term performance potential and stability across envi-ronments for nine apple rootstocks tested in the 1980–81 NC 140 trial. Fruit Varieties Journal 45, 208–212.

Olien, W.C., Ferree, D.C., Bishop, B.L. and Bridges, W.C., Jr (1995) Prediction of site index and apple root-stock performance from environmental variables. Fruit Varieties Journal 49, 179–189.

Parnia, P., Dutu, I., Vlãdianu, D., Movileanu, M., Mazilu, C., Bragãu, A. and Stefan, D. (1997) 30 de Ani de Activitate (1967–1997). Institutul de Cercetare, Desvoitare Pentru Pomicoltura Pitesti, Pitesti-Mracineni, pp. 45–52.

Parry, M.S. (1965) Field trials of Malling–Merton rootstocks. In: Annual Report of East Malling Research Station for 1964. Maidstone, UK, pp. 91–96.

Parry, M.S. (1976) Regulation of growth and cropping: rootstocks: apple rootstocks: height of budding.

In: Report of East Malling Research Station for 1975. Maidstone, UK, p. 35.

Parry, M.S. and Rogers, W.S. (1968) Dwarfing interstocks: their effect on the field performance and anchorage of apple trees. Journal of Horticultural Science 43, 133–146.

Parry, M.S. and Rogers, W.S. (1972) Effects of interstock length and vigour on the field performance of Cox’s Orange Pippin apples. Journal of Horticultural Science 47, 97–105.

Pearl, R.T. (1932) Apple rootstocks I–XVI. Journal of the South-Eastern Agricultural College, Wye, Kent 30, 194–213.

Perry, R.L. (1990) Mark in the rootstock arsenal. Compact Fruit Tree 23, 1–3.

Perry, R.L. (1992) The relative susceptibility of rootstocks to fire blight. Proceedings of the Michigan State Horticultural Society 122, 72–75.

Pieniazek, S.A., Czynczyk, A. and Zagaja, S.W. (1976) Apple rootstocks from other socialist countries evaluated in Poland. Compact Fruit Tree 9, 52–57.

Preston, A.P. (1954) Apple rootstock studies: the MIX crosses. In: Annual Report of East Malling Research Station for 1953. Maidstone, UK, pp. 88–94.

Preston, A.P. (1955) Apple rootstock studies: Malling Merton rootstocks. Journal of Horticultural Science 30, 25–33.

Preston, A.P. (1966) Apple rootstock studies: fifteen years’ results with Malling–Merton clones. Journal of Horticultural Science 41, 349–360.

Preston, A.P. (1970) Apple rootstock studies: growth and cropping of trees on M.26, M.VII and M.II. In:

Annual Report of East Malling Research Station for 1969. Maidstone, UK, pp. 99–101.

Preston, A.P. (1971) Apple rootstock 3431 (M.27). In: Annual Report of East Malling Research Station for 1970. Maidstone, UK, pp. 143–147.

Privé, J.-P. and Embree, Ch. (1997) Low temperature injury and recovery of apple rootstocks. Acta Horticulturae 451, 179–186.

Proctor, J.T.A., Hutchinson, A. and Pierce, W.F. (1974) A 10-year trial of seven apple cultivars on Malling 26 rootstock. Canadian Journal of Plant Science 54, 661–665.

Quamme, H.A. (1990) Cold hardiness of apple rootstocks. Compact Fruit Tree 23, 11–15.

Quamme, H.A. and Brownlee, R.T. (1990) Rootstocks for high density apple plantings in British Columbia.

Technical Report 90-02, Agriculture Canada, Summerland, British Columbia.

Quamme, H.A., Hampson, C.R. and Brownlee, R.T. (1999) Apple rootstock evaluation for the climate of British Columbia, Canada. In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 87–88.

Robinson, T.L., Cummins, J.N., Hoying, S.A., Johnson, W.C., Aldwinckle, H.S., Norelli, J.L., Momol, M.T.

and Saygili, H. (1999) Orchard performance of fire blight-resistant Geneva apple rootstocks. Acta Horticulturae 489, 287–294.

Rogers, W.S. (1958) Malling 26, a new semi-dwarfing apple rootstock. In: Annual Report of East Malling Research Station for 1957. Maidstone, UK, pp. 48–49.

Rom, C.R., Rom, R.C. and Stasiak, M.J. (1990) Size controlling apple rootstocks affect growth, spur qual-ity, foliar nutrition and productivity. Compact Fruit Tree 23, 17–21.

Sadowski, A. and Hrotko, K. (1999) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, 131 pp.

Sax, K. (1949) The use of Malus species as rootstocks for apple. Proceedings of the American Society for Horticultural Science 53, 219–220.

Schmidt, H. (1988) Criteria and procedures for evaluating apomictic rootstocks for apple. HortScience 23, 104–107.

Schupp, J.R. and Ferree, D.C. (1987) Effect of root pruning at different growth stages on growth and fruit-ing of apple trees. HortScience 22, 387–390.

Soejima, J., Bessho, H., Tsuchiya, S., Komori, S., Abe, K. and Kotoda, N. (1998) Breeding Fuji and perfor-mance on JM rootstocks. Compact Fruit Tree 31, 22–24.

Soumelidou, K., Morris, D.A., Battey, N.H., Barnett, J.R. and John, P. (1994) Auxin transport capacity in relation to the dwarfing effect of apple rootstocks. Journal of Horticultural Science 69, 719–725.

Stover, E.W. and Walsh, C.S. (1994) Tissue proliferation on Mark rootstock shanks: effect on tree growth and correlation with site factors. HortScience 29, 475.

Trajkovski, V. and Andersson, G. (1980) [Rootstock breeding]. In: Report for 1978–79. Division of Fruit Breeding, Swedish University of Agricultural Science, Balsgård, pp. 36–48.

Tubbs, F.R. (1972) Research fields in the interaction of rootstocks and scions in woody perennials, Parts 1 and 2. HortAbstracts 43, 247–253, 325–335.

Tubbs, F.R. (1980) Growth relations of rootstocks and scion in apples. Journal of Horticultural Science 55, 181–189.

Tydeman, H.M. (1933) Breeding experiments with ‘Paradise’ apple rootstocks. Journal of Pomology 11, 214–216.

Tydeman, H.M. (1943) Further studies on new varieties of apple rootstocks. Journal of Pomology 20, 116–126.

Tydeman, H.M. (1953) A description and classification of the Malling Merton and M.XXV apple root-stocks. In: Report of East Malling Research Station for 1952. Maidstone, UK, pp. 55–63.

Tydeman, H.M. (1955) Descriptions of the Malling apple rootstocks. In: Report of East Malling Research Station for 1954. Maidstone, UK, pp. 64–66.

Van Oosten, H.G. (1977) Growth and yield of Golden Delicious on M.9A and three other M.9 rootstock sources. Acta Horticulturae 75, 157–161.

Van Oosten, H.G. (1986) Intraclonal variation in M.9 rootstock and its effects on nursery and orchard per-formance. Acta Horticulturae 160, 67–78.

Verzilin, A.V., Verzilina, N.V. and Kuzin, A.I. (1999) ‘Malysh Budagovskogo’ is the new dwarfing apple rootstock. In: Sadowski, A. and Hrotko, K. (eds) Apple Rootstocks for Intensive Orchards. Proceedings of an International Seminar, Warsaw, 18–21 August 1999. Department of Pomology, Warsaw Agricultural University, Poland, and Department of Pomology, University of Horticulture and Food Industry, Budapest, Hungary, pp. 115–116.

Warner, J. (1991) Rootstock affects primary scaffold branch crotch angles of apple trees. HortScience 26, 1266–1267.

Webster, A.D. (1994) Rootstock and interstock effects on deciduous fruit tree growth and cropping: a brief review. Compact Fruit Tree 27, 5–16.

Webster, A.D. (1995) Temperate fruit tree rootstock propagation. New Zealand Journal of Crop and Horticultural Science 23, 355–372.

Webster, A.D. and Hollands, M.S. (1999a) Apple rootstock studies: comparison of Polish, Russian, USA and UK selections as rootstocks for the apple cultivar Cox’s Orange Pippin (Malus domestica Borkh.).

Journal of Horticultural Science and Biotechnology 74, 367–374.

Webster, A.D. and Hollands, M. (1999b) Orchard comparisons of ‘Cox’s Orange Pippin’ grown on selec-tions of the apple rootstock M.9. Journal of Horticultural Science and Biotechnology 74, 513–521.

Webster, A.D. and Tobutt, K. (2001) Breeding and selection of new apple rootstocks at Horticulture Research International-East Malling. Proceedings of the 7th International Symposium on Orchard and Plantation Systems, Nelson, New Zealand. Acta Horticulturae 557, 189–192.

Webster, A.D., Smith, R.A. and Watkins, R. (1986) Apple rootstock studies. I Preliminary evaluations of several MM.106 × M.27 hybrids. Journal of Horticultural Science 61, 429–437.

Webster, A.D., Atkinson, C.J., Lucas, A.S., Vaughan, S.P. and Taylor, L. (2000) Interactions between root restriction, irrigation and rootstock treatments on the growth and cropping of ‘Queen Cox’ apple trees: effects on orchard growth and cropping. Journal of Horticultural Science and Biotechnology 75, 181–189.

Webster, C.A. and Jones, O.P. (1989) Micropropagation of the apple rootstock M.9: effect of sustained sub-culture on apparent rejuvenation in vitro. Journal of Horticultural Science 64, 421–428.

Welander, M. (1998) The use of genetic engineering in fruit tree improvement. Currents Uppsala 17–18, 47–50.

Welander, M. and Zhu, L.H. (2000) The rooting ability of rolB transformed clones of the apple rootstock M26 and its relation to gene expression. Acta Horticulturae 521, 133–138.

Wertheim, S.J. (1985) Vorstschade bij appel en peer. Verschillen tussen rassen, onder- en tussenstammen.

Fruitteelt 75, 1376–1380.

Wertheim, S.J. (1991) Onderzoek naar onderstammen volop in beweging. Fruiteelt 81(15), 14–15.

Wertheim, S.J. (1992) M.20, een appelonderstam voor de toekomst. Fruiteelt 3, 11.

Wertheim, S.J. (1997) Useful differences in growth vigour between sub-clones of the apple rootstock M.9.

Acta Horticulturae 451, 121–128.

Wertheim, S.J. (1998) Apple rootstocks. In: Rootstock Guide: Apple, Pear, Cherry, European Plum. Fruit Research Station, Wilhelminadorp, The Netherlands. pp. 19–59.

Wertheim, S.J. and Callesen, O. (2000) Results of multi-site interstem trials with apple.

Gartenbauwissenschaft 65(6), 251–259.

Wertheim, S.J. and Kunneman, B.P.A.M. (1993) Onderstammen uit weefselkweek zijn bruikbaar. Fruitteelt 83(21), 20–21.

Zagaja, S.W. (1980) Performance of two apple cultivars on P. series dwarf rootstocks. Acta Horticulturae 114, 162–169.

Zagaja, S.W., Czynczyk, A., Jakubowski, T. and Omiecinska, B. (1988) Breeding and evaluation apple rootstocks for northern Europe. HortScience 23, 109–112.

Zagaja, S., Jakubowski, T. and Przybyla, A. (1991) P.60 a new vegetative apple rootstock. Fruit Science Reports 18, 25–30.

S.J. Wertheim

1

and Anthony D. Webster

2

1Fruit Research Station, Randwijk, The Netherlands; 2Crop Science Department, Horticulture Research International, East Malling, West Malling, Kent, UK

6.1 Introduction 126

6.2 Seed Propagation 126

6.3 Vegetative or Clonal Propagation 127

6.3.1 Stooling 128

6.3.2 Layering 129

6.3.3 Cuttings 131

6.4 Micropropagation 135

6.4.1 Propagation in vitro 135

6.4.2 Practical applicability 136

6.5 Tree Raising 136

6.5.1 Site choice 136

6.5.2 Planting distance 137

6.5.3 Rootstock cutting (heading) back and bleeding 137

6.5.4 Plant material 138

6.5.5 Tree support 139

6.5.6 Trunk cleaning 139

6.5.7 Other types of plant material 140

6.6 Budding and Grafting 140

6.6.1 Bud wood 141

6.6.2 Rootstocks, budding height and site 141

6.6.3 Budding and grafting methods 141

6.6.4 Tying and after care 142

6.6.5 Bench grafting 143

6.7 Branching 143

6.7.1 Manipulation of branching by hand 143

6.7.2 Manipulation of branching by chemicals 144

6.8 Defoliation and Digging Up (Lifting) Trees 146

© CAB International 2003. Apples: Botany, Production and Uses

(eds D.C. Ferree and I.J. Warrington) 125

6.1 Introduction

In nature, apple trees multiply by seeds, but when apple trees are grown commercially or by the home gardener the common propaga-tion method is to bud or graft scion cultivars on to selected rootstocks. The reason for propagation by means of rootstocks is that apple cultivars are not true to type when propagated by seed and, at least in former days, difficult, if not impossible, to propagate by vegetative means. Thus, when a seedling with good fruiting characteristics was found, it could not be multiplied by sowing its seeds or by use of layering or cutting techniques.

Today, vegetative propagation techniques have improved so much that scion cultivars can be multiplied by cuttings or by micro-propagation in the laboratory. Nevertheless, the use of rootstocks has remained dominant, because most of the currently grown scion cultivars, once self-rooted, do not perform well in the orchard. Scion trees propagated

‘on their own roots’ are more vigorous, come later into bearing and crop less efficiently than those on good rootstocks.

Apple seedlings must have been the first rootstocks used extensively by early horticul-turists, largely because seeds were readily available. Only after it was observed that some apple cultivars could be easily propa-gated by layering did vegetative or clonal

Apple seedlings must have been the first rootstocks used extensively by early horticul-turists, largely because seeds were readily available. Only after it was observed that some apple cultivars could be easily propa-gated by layering did vegetative or clonal

In document Apples - Botany, Production and Uses (Page 132-139)