MemoirsoftheMuseumofVictoria 50(2): 379-402 (1990) ISSN0814-1827
ACCALATHURA (CRUSTACEA: ISOPODA: PARANTHURIDAE) FROM NORTHERN AUSTRALIAAND ADJACENT SEAS
By GaryC. B. Poore and HelenM. Lew Ton
Departmentof Crustacea, Museum ofVictoria, SwanstonStreet Melbourne, Victoria 3000, Australia
Abstract
Poore, G.C.B. andLewTon, H.M., 1990.Accalathura(Crustacea: Isopoda:Paranthuri- dae)fromnorthernAustraliaandadjacent seas.MemoirsoftheMuseumofVictoria50(2):
379-402.
Elevennewspeciesof Accalathura Barnardfromreefandshelfenvironmentsinnorthern AustraliaandtheCoral Seaare figuredanddescribed.Most are typicalofthegenusinthe possession ofnarrowuropodal exopods:(A.avena, A.eulalia,A. poa, A. spathia, A.themeda, A. triodea, A. vulpiaandA. zoisia);butthreedifferfromallothers inhavingaverybroad exopod:(A.dimeria, A.oryzaandA.sehima).Accalathura barnardi(Nierstrasz)fromIndon- esiaisredescribed andakey tothe speciesfrom theregionis presented. Speciesfromthe IndianOcean, which haveoften been misidentified, are discussed.
Introduction
Thetropicalanthurideanisopodfauna ofAus-
tralia is dominated by species of the anthurid genus Amakusanthura Nunomura (Poore and Lew Ton, 1988) and the paranthurid genus Accalathura Barnard. In this contribution new
species ofAccalathura from this region are de- scribed. The genus has been recorded several times from the Indian Ocean and south-east Asia. These records are reviewed.
The genus is distinguished from all other paranthuridsbythepossessionofa multiarticu- late flagellum on both pairs of antennae. Most
species are inhabitants ofsoft sandy or muddy
sublittoral substrates but some are recorded fromcoral debris.Thegenusincludesthelongest anthuridean known, A. gigantissima Kussakin fromtheSouthernOcean(Poore, 1981:Wagele, 1985).Tropical species,incontrast, areofasize
more typical of anthurideans generally.
Indo-west-Pacific species ofAccalathura
Thesenewspecies bring to22 the number of speciesofAccalathura described. Two, possibly four, are known from the West Atlantic, one fromtheNorth Pacific,twofromsouthernAus-
tralia,and one fromAntarcticseas(Poore, 1980, 1981).
Seven species of Accalathura have been de- scribed or recordedfrom the Indian Ocean and south-east Asia but a review ofpublished de- scriptions and figures suggests that some are
misidentifications. A complete re-examination
ofall the material is beyond the scope ofthis paper.
The most widely reported Indian Ocean
species is A. borradailei (Stebbing, 1904) origi- nally described from the Maldives. It has a roundedtelson anda relatively broaduropodal exopod. Recordsfrom India (Chilton, 1924;Pil- lai, 1966)fitthisdescriptionbut thosefromeast Africa(Monod, 1972a,b)areofanotherspecies with anacutetelson and narrowexopod. Accal- athura laevitelson (Kensley, 1975), described fromamanca,issimilarinallcriticalcharacters and may be an appropriate name for this east African species. These records may be of the same species which Kensley (1980, 1988) reported from thesamearea and from Aldabra Atoll and which he referred to A. sladeni (Steb- bing, 1910).
Accalathurasladeni wasdescribed from Car- gadosCarajos,northof Mauritius,andalthough
it is similar to the material figured by Monod
and Kensley (1980) there are subtle differences.
Inparticular,theappendixmasculina ofA. slad- eniis simple whilethat of Kensley's male hasa bifid apex. It seems,therefore, that at leasttwo similar speciesfromtheIndianOceanhave been referred to A. sladeniandthatbothare different from A. borradailei. The name A. sladeni was alsousedby Hale(1937)foran Accalathurafrom South Australia but this material has subse- quentlybeendescribed asA. bassiPoore, 1981.
Aspeciessimilar to theAfricanone,withabifid
appendix masculina, was recorded from sou-
379
https://doi.org/10.24199/j.mmv.1990.50.11
31 March 1990
them Western Australiaby Thomson(1951)as A. gigas whichit is not (Poore, 1981).
ThetypesofthreeIndonesianspeciesfromthe
"Siboga"collections(Katanthura barnardi Nier- strasz, 1941, Metanthura indica Nierstrasz, 1941 and M. normani Nierstrasz, 1941) were examinedbyGCBP inAmsterdam in 1981. All
have been placed inAccalathura (Poore. 1980).
The types are not available for detailed illus- tration and onlyA. barnardi isre-illustrated in this contribution from new material. Accala- thura indica is similar to A. sladeniinthe pos- sessionofa digitiformpalmarlobeon pereopod
1 (not triangular as figuredbyNierstrasz).Kens-
ley(1977, 1982) recordedthis speciesfromeast Africa but his illustrations suggest the same
species as thathecalledA.sladeniin 1980.Accal- athura normani (Nierstrasz, 1941), described froma male,is notsufficientlywelldescribedto be easilyrecognised.
In summary, A. borradailei is definitely recorded fromthe Maldives and India, A. slad- enifrom Cargados Carajos,andA. barnardi, A.
indicaandA normanifrom Indonesia. Thereis
certainly at least one other species from East Africaand AldabraAtoll (variously reportedas A. borradailei, A. sladeni, andA. indica)which maybethesameas A. laevitelsondescribedfrom that area. The identity of a WesternAustralian speciesis also stillenigmatic.
SpeciesfromAustralia andadjacent seas
Many speciesofAccalathura are superficially quite similarwhichaccountsforthe misidentifi- cationofseveralspeciesfromtheIndian Ocean.
Butin thiscontributionanewgroupofspeciesis
reported (A. dimeria, A. oryza, A. sehima) in
which a very broad uropodal exopod is held erectoverthe telson;inallotherspecies theexo- pod is linear. Accalathura dimeria differs from theothertwoinhavingarounded telsonicapex and taperingpropodus on pereopod2.A. oryza
is much smaller than A. sehima and differs in
pereopod 2 and uropodal endopod.
Theremainingspecies areverysimilarandare not easily gouped on morphological criteria.
One group ofspecies is those in which the pal-
marlobe ofthe propodusofpereopod 1 is well defined: A. avena, A. indica, A. poa, A. triodea andA. vulpia. In the remainingspecies thepal-
mar lobe is poorly defined.
Two species (A. avena and A. vulpia) differ
fromthe others inhavingthe uropodalexopod moderately broad (2.4 to 3 times as long as broad)andnotreachingtheendofthepeduncle.
In most speciestheuropodalexopod isabout 4
times as long as wide and exceeds the ped- uncle.
Thetelsonisusuallyabout2.5timesaslongas widebutinthreespecies(A.themeda,A.poaand A. zoisia) it ismuch narrower,about 3times as longas wide. The apex istypically acute butA.
vulpia differs from the others in having a roundedapex,similar in some waystothe three speciesofthe broad-exopod groupin whichthe telson apex is very obtusely angled.
Severalspeciesare dorsallyanddorsolateral^
pigmentedfrom the headto telson: A. avena,A.
dimeria,A. eulalia, A.poa andA. themeda. Pre- served material ofthe other species is not pig-
mented but the possibility of pigment having been lost cannot be discounted.
Similaritiesbetweenspeciesarenot discussed further in this paper.
Methods
Inthis contribution the new species,Accala- thura themeda, isfiguredand describedindetail first.Exceptforthe three specieswithbroaduro- podalexopods.there areonly minordifferences between this species and the rest in the shape and setation ofthe antennae, mouthparts, pos- teriorpereopods. and pleopods. Forthese con- servative characters A. themedamaybe takenas typical of the genus (Figs 1-3). Other species have been dissected andexamined indetailbut only the most species-diagnostic features are figured: tailfan.telson.uropods and first,second andfourthpereopods.Alllimbsarefromtheleft side and only distal articles are drawn. Illus- trations are somewhat simplified. Forpereopod
1. setae and the bases ofthe palmar spines are shown; the spines on and near the palmarlobe are drawn at a higher magnification; and the lateral setal row is not figured. On other parts onlythebases ofmanysetaeare figuredandlong setae, e.g.. on the telson and uropod have been truncated. The written diagnoses are similarly abbreviatedtoconcentrateon importantcharac-
ters. When known, male characters are des- cribed.
In figures the following abbreviations are used: Al. 4.2, antennae 1, 2; MD. mandible;
MDp, mandibular palp; MX, maxilla; MP,
maxilliped: P1-P7, pereopods 1-7; PL1-PL5, pleopods 1-5; T, telson; UN, UX, uropodal endopodand exopod;AM,appendixmasculina.
Figures marked a or unmarked are of the holotype; those marked b are from a male paratype.
Following a pattern established earlier, the new species arenamed forgenera of Australian plants; this timeall are genera ofgrasses.
ACCALATHURA(ISOPODA) FROMTROPICAL AUSTRALIA 381
Material islodged intheMuseumofVictoria, uropod.The endopodmaywellexceedthe telson Melbourne (NMV), Australian Museum, Syd- not barely exceed as previously stated) and the ney (AM). Queensland Museum, Brisbane exopod maybebroad (not always narrow).The (QM), NorthernTerritoryMuseumandArt Gal- multiarticulate antennal flagella are unique
lery, Darwin (NTM), Zoological Museum, withintheAnthuridea and areaprobablesyna-
Amsterdam (ZMA), and Zoological Museum, pomorphy.
Copenhagen (ZMC). In males of Accalathura, as in many other anthurideangenera, the pleon, uropodsandtel- Accalathura Barnard sonaremoreelongatethaninfemalesandjuven- Typespecies. Calathura cremdata Richardson,
j
les
- The
f
anteima
' flagellum consists of two
qq^ types ot articles; proximal, discoid articles
whichbeardense whorls ofaesthetascs,anddis-
Remarks. Poore (1980, 1981) provided a talcylindricalarticleswhich donot.Pereopod 1
synonymy, diagnosis and description of this is moreelongate and denselysetose, butpereo- easilyrecognisedgenus.Thediagnosismustnow pods2 and 3 mayalso be modified with strong beexpandedtoaccommodate variability inthe teeth.
Key to speciesofAccalathura from northernAustralia andadjacent seas
1. Uropodal exopod not more than twice as long as broad, with a well-
developeddorsallobe,not reachingtoendof peduncle;pereopod 1 with
palmarlobe defined by a right angle 2
— Uropodal exopodmorethan2.4timesaslongasbroad,withouta dorsal lobe, reaching to endofpeduncle or not; pereopod 1 with palmarlobe
defined bya right angleor obtuse angle 4
2. Uropodal endopod exceeding end oftelson; apex of telson obtusely
angled • 3
— Uropodal endopod reachingas far as telson; apexoftelson rounded . A. dimeria
3. Pereopod 2 propodus oval (twice as long as wide); uropodal endopod
withgently convex inner margin A. oryza
— Pereopod2propoduslinear(3timesaslongas wide);uropodalendopod
with strongly convex innermargin A, sehitna
4. Pereopod 1 propodal palmarlobe right-angled 5
— Pereopod 1 propodal palmarlobe obtusely-angled 9
5. Uropodal exopodnotreachingtoendof peduncle; peduncle innerdistal
angle half as longas endopod A. avena
— Uropodal exopodreachingbeyondoras farasendof peduncle; peduncle inner distal angle lessthan halfaslong asendopod 6 6. Uropodal exopod 2.4times as long as wide; endopod as long as wide;
apex oftelson rounded A. vulpia
— Uropodal exopod about 4 times as longas wide; endopod longerthan
wide; apex oftelson acute 7
7. Pereopod 1 palmar lobe broadly triangular, near midpoint ofoverall length of propodus; uropodal endopod parallel-sided, with broadly
angled apex A- triodea
Pereopod 1 palmarlobedigitiform,aboutone-thirdalonglengthofpro- podus; uropodal endopod tapering, elongate, triangular 8 8. Uropodal exopod 4.5 timesas longas wide; peduncle with acute inner
angle, one-third aslongas endopod A. poa
— Uropodal exopod 3.7 timesas long as wide; peduncle with blunt inner
angle, one-tenth as longas endopod A. indica
9 Telson taperingfrom nearbase;uropodalendopodwith2 setaeoninner
margin
.A.
themeda— Telson widest near midpoint; uropodalendopodwith3ormoresetaeon
inner margin 10
382 G. C. B POOREAND H. M. LEWTON
10. Tclson apexacute *
— Telson apex rounded '^
1 Uropodalendopodapexacute;pereopod2propodus2.4timesaslongas
wide A- eulalia
— Uropodalendopodapexrounded;pereopod 2propodus2.0timesaslong
as wide A- barnardi
12. Uropodalendopod morethan twiceaslongaswide;male appendix mas- culina with 2 distal fingers; male pereopod 2 propodus with slightly
convex palm 4- spathia
— Uropodal endopod lessthan twiceas longaswide; male appendix mas- culina with 3 distal fingers; male pereopod 2 propodus with strong
proximal palmar lobe 4. zoisia
. .. ., . with proximal palmarlobe separatedfrom palm Accalathura themeda sp. nov.
by
^^ ^ Qf,w percopod 2 propodus
Figures 1-3 2-7timesaslongaswide, withconvex palmbear- ing 8 spines on distal two-thirds. Pereopod 4 Materialexamined. 1 male. 1 female,44juveniles; 5- carpusand propoduswith 5and 6spinesrespec- 14"n
!
11 ..(,,„, . -. ., , ,,. . tivelv; propodus 4.5 limes as long as wide.
Holotype. Coral Sea (French Territory), Chester- - h field Reefs Long Is. ('f52'S,158M9'E) seaward
fl
. .
Hcad eon and ,eon wkh .
edge 12m,NX. Bruce,5May1979,QMW8130 with
^/^
an[j dorsolatcrai pigment paUern.2 slides, luvenile, 11.4mm. , .. , ,. /" .* . -,
Paratypes, Typelocality,QM W8I21(I specimen), Ratio ofdorsal lengths of head, pereonites 1-7, W8126(2), W8128(2), NMV J10125(8). Chesterfield plcon, telson-0.6: 1.0: 1.1 : 1.1 : 1.0: 1.0: 1.1 :
Reefs,LongIs.,seawardedge, 15m,N.L.Bruce, 6May 0.4 : 0.7 : 0.8.
1979,QMW15989(1).Bennett Is.: inner reef edge, 12 Head with short rostrum; eyes pigmented, m, N.L. Bruce, 6 May 1979, QM W8104(3); lagoon, Articulationsofpcrcopodsmarkedbygroups of rearward edge, 1 m, N.L. Bruce. 7 May 1979. long setae. Pleonites free. 1-5 of equal length,
W8124(4);Nend oflagoon, 1 m, N.L. Bruce, 8 May
pleonite 6 longer and fused t0teison but wjth a
1979,VV8101(2juveniles, 1female).CayNofLongls.
„ k d transverSe ridge dorsallv between seaward slopes, 15 m, N.L. Bruce, 8 May 1979, QM . .
b
WKKP(l) the two.
Australian Coral Sea Territory. Magdalaine Cay Antenna 1,2.3timesaslongashead;peduncle
(16°37'S, 150°17'E): beach rock, N.L. Bruce, 26 Apr with second article shortest, tlagellum longer
1979,QMW8098(2);deadcoral, 10m,N.L.Bruce,27 than peduncle, ofbasalarticle plus 17isometric Apr 1979, W8122(l). Mellish Reef (I7°25'S, articlesofwhich4lo 14each bears 1 aesthetasc.
155°50'E): reefedge-dropoft',13m,N.L.Bruce, 2May Antenna 2, 3.5 times aslong as head; flagellum 1979,QM W8097(2); back reefbommie. 10m, N.L. shorterthan peduncle, of27 setosearticles.
Bruce. 1 May 1979 W8123(1); lagoon, 10 m N.L. Mouthparts produced well forward bevond
Bruce, May 979, W8 25(3); back reeledge. 20 m. , , .„ ... , , ... ., .. , 'c ,
N.L. Bruce. 1 May 1979, W8127(l). Marion Reef cyelobes Mandibular palp with 3 articles fir t
(19'10'S, 152M7'E), lagoonpinnacle,2m,N.L.Bruce. short wltn ' seta
- 8econd w,th 3 setae- third tal'
13 May 1979, QM W8120(2). Marion Reef, Brodie catewithlongitudinalrowol 16evensetae plus1
Cay, reeffront. 15-20 m, N.L. Bruce, 12 May 1979, longer seta. Maxilla 1 a finely serrate spine.
QM W8096 with 1 slide (1 male, 12.4 mm), NMV Maxilliped with small epipod, coxa and basis
J10124(2). fused tohead, enditeabroad blade withsubter-
Othermaterial.Qld.YongeReef(14°37'S,15°38'E). minai seta; paip wjth articles 1 and 2 fused
2 m, P. Hutclnngsand P.Weaie, 19Jan 1975(stn 75 (togetherwith 5ventralsetae),articles3-5 fused
[fvwk »M« ? )-,o*Tk,n S
/
3 ^ lo • (together with 1 dorsal and 14apical setae).
145 28E), B. Kensley,Jan 1982, NMV J12853(2). °
. , ... , . . • ,• ,- ,
Pereopod 1 with basis and ischium of equal Diagnosis. Dorsallypigmented. Telson tapering lengths, merus completely enclosing carpus;
from near base to an acute apex, 3.0 times as propodus with proximal palmar lobe separated longaswide. Uropodal peduncle reaching 85% from palm by rounded angle of c. 130°, palm of length of telson, distally defined by acute withamesialrowofc.30setae,alateralrowofc.
angles;endopoddistally rounded, exceedingtel- 60 serratespines of variouslengths, lateral face son by half its length; exopod tapering from with row of 15 setae. Pereopod 2 with setose base, 4.3 timesas longas basal width, reaching margins on basis and ischium; propodus 2.7 just beyond peduncle. Pereopod 1 propodus times on long as greatest width, palm convex.
ACCALATHURA (ISOPODA)FROMTROPICAL AUSTRALIA
Figure 1.Accalathura themeda. Holotype, 11.4mm, QMW8130.
Figure 2.Accalathura themeda. Holotype, 11.4mm,QM W8130.
ACCALATHURA (ISOPODA)FROM TROPICAL AUSTRALIA 385
Figure3.Accalathurathemeda.a,holotype, 11.4mm,QMW8130.Antenna 1peduncle, 6 basaland4terminal flagellar articles;antenna2 peduncleand2flagellar articles;maxilliped withdetailofmesial apex,b,paratype male, 12.4 mm, QM W8096.
withlateraland mesial setaeand 8 submarginal spines (distal ones complex) on mesial face.
Pereopod 3 similar to 2, propodus more elon- gate. Pereopods 4-7 becoming longer poster- iorly; basis, ischium and merus setose; carpus 3-4timesaslongaswide(narrowerposteriorly), with4-5marginalspines;propodus 4-5timesas longaswide, with6-7 marginalspines;dactylus about halflength ofpropodus.
Pleopod 1 exopod,2.2timesaslongasbroad, with 23 marginal plumose setae; cndopod
shorter,with4 terminalplumosesetae.Pleopods 2-5 shorterthan pleopod 1, similar, with rami similar,endopodwith 5setae,exopodwith9-12
setae.
Uropodal pedunclereaching85%oflengthof telson, distally defined by acute angles sur- rounding endopod; endopod distally rounded, exceedingtelson byhalfitslength,with 2mesial setae and dense setation distally and laterodis- tally,andwith 3separatebrush-setaeplusgroup of3 brush-setae dorsally; exopod reachingjust beyond peduncle, tapering frombase, 4.3 times aslongas basalwidth, marginallysetose, mostly simplesetae dorsally,plumoseventrally.Telson tapering from near base to an acute apex, 3.0 timesas longas wide, apex withc. 18 long sub- marginal setae plus pair of small setae at apex.
Male. Pereon, pleon and pereopods more elongatethanjuvenile.Antenna I flagellumwith 23 articles each with ring ofnumerous aesthe- tascs, plus 6 narrow terminal articles without aesthetascs.Pereopod 1 propodusdenselysetose mesially. Pereopod 2 propodus narrower dis- tally than in juvenile, with marginal row of 8 spines.
Distribution. Coral Seaand northernGreat Bar- rier Reef; coral rubble at 1-20 mdepth.
Accalathuraavena sp. nov.
Figure 4
Materialexamined. 10 juveniles, 12.0-28.7 mm.
Holotype. Old, N of Magnetic Is. (19°08'S, I46°50'E),7m,P.Arnold, 25Aug1976,NMVJ10101 with I slide, juvenile. 28.7mm.
Paratypes. Qld (all collected by P. Arnold in
Townsvilleregion). BowlingGreen Bay,muddysand,
l7m,7Aug1975,NMVJI0103(1specimen);mud,13 m,9 Dec 1975, NMV.110107(1);sand, 18 m, 16Apr
!975,NMVJ10105(l);mud, 14m, 17Jun1975,NMV J10106O). Halifax Bay, 15 m, 24 May 1976, NMV
J10102(1); muddy sand, 13 m, 24 Feb 1971, AM
P39440O), NMV.110108(1), QM W15990(l).Cleve- land Bay, mud,3 m, 4Jun 1974NMVJ101 04(1).
Diagnosis. Dorsally pigmented. Telson widest two-thirds along, tapering to a broadly acute
apex, 2.5 times aslongas wide. Uropodal ped- uncle reaching 95%oflength oftelson, mediod-
istally definedbyabroad triangularlobehalf as longasendopod;endopod distally tapering, 1.5 timesas longas wide, exceedingtelson byone- thirdofitslength;exopodwidestatmidpoint3.0 timesaslongaswide, not reaching base of endo- pod. Pereopod 1 propoduswith strongproximal palmarlobe,separatedfrompalmbyrightangle.
Pereopod 2propodus2.7 timesaslongaswide, distally narrower, with convex palm bearing 8 spines. Pereopod 4carpus and propodus with 5
and 4 spines respectively, 3.8 times as longas wide.
Distribution. Queensland (type locality only), shallow shelfnear Townsville. 7-18 m.
Accalathura barnardi (Nierstrasz) Figure 5
Katanthura barnardi Nierstrasz. 1941: 243-247,
figs 1-13.
Accalathura barnardi.—Poore, 1980: 59.
Material examined. I male,1 female,1juvenile; 11.7- 15.1 mm.
Holotvpe.Indonesia, SoloStrait. 113 m,M.Weber,
8 Feb 1900 ("Siboga" Expedition stn 305), ZMA
Is.100.620, female, 14mm.
Other material. Indonesia. S of Bali (8°46'S,
II5°15'E),coralsand. I9m. 12 Sep 1951 ("Galathea"
stn483).ZMC (juvenile 15.1 mm, male 11.7mm).
Diagnosis. Dorsal pigment possible. Telson widest at midpoint, lateral margins evenly curved to moderately acute apex, 2.5 times as longas wide. Uropodal peduncle reaching 90%
oflength oftelson, distally defined by a broad triangular mesial projection; endopod distally rounded, exceedingtelsonbyhalfitslength;exo- podtapering overmostofitslength,4.0timesas long as wide, reaching just beyond peduncle.
Pereopod 1propoduswithproximal palmarlobe separated from palm by a obtuse angle. Pereo- pod 2 propodus2.1 timesaslongaswide,ovate, with 9 marginal spines. Pereopod 4 carpus and propodus with 6 and 8 spines respectively;
propodus 3.5 times aslong aswide.
Male. Pereopod 2 propodus palm irregular.
Appendix masculina with elongate subapical blade.
Distribution. Indonesia, 19-113 m.
Accalathura dimeria sp. nov.
Figure 6
Material examined.2sub-males, 1female,2juveniles;
10.3-13.7 mm.
ACCALATHURA(ISOPODA)FROM TROPICAL AUSTRALIA 387
Figure4. Accalathuraavena. Holotype,28.7mm, NMV J10101.
Holotype. Qld,HalifaxBay(19°05'S, 146°43'E), 10 m, P. Arnold, 24 Aug 1976 (TBS stn), NMV
J10109(juvenile, 11.6 mm).
Paratypes.Qld. Halifax Bay, 3-13 m,coarsesilt to very fine sand, P. Arnold, various dates (TBS stns),
NMVJ10110 with 2 slides (1 specimen), JlOlll(l), Jl0112(1 sub-male), J10113(1 sub-male), QM
W1599K1).
Diagnosis. Dorsally pigmented. Telson almost
parallel-sided formuch oflength, tapering to a broadlyroundedapex, 2.4timesaslongaswide.
Uropodalpeduncle reaching abouttwo-thirdsof length oftelson, distallydefined by acute angle mesially and broad lobe laterally; endopod narrow,apically subacute,reachingtoendoftel- son; exopod strongly dorsally lobed, about 1.5 times as long as greatest width, ventral lobe reaching three-quarters along peduncle. Pereo-
Figure 5. Accalathura barnardi. a,juvenile, 15.1 mm, ZMC. b, male, 11.7 mm, ZMC.
