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Vol. 23, No. 4 (2000) 261–270 S0161171200001010 © Hindawi Publishing Corp.

STABILITY OF THE POSITIVE STEADY-STATE SOLUTIONS

OF SYSTEMS OF NONLINEAR VOLTERRA DIFFERENCE

EQUATIONS OF POPULATION MODELS WITH

DIFFUSION AND INFINITE DELAY

B. SHI

(Received19 June 1998)

Abstract.An open problem given by Kocic andLadas in 1993 is generalizedandconsid-ered. A sufficient condition is obtained for each solution to tend to the positive steady-state solution of the systems of nonlinear Volterra difference equations of population models with diffusion andinfinite delays by using the methodof lower andupper solutions and monotone iterative techniques.

Keywords and phrases. Nonlinear Volterra difference equations, diffusion, infinite delays, positive steady-state solutions, methods of lower and upper solutions, monotone iterative techniques.

2000 Mathematics Subject Classification. Primary 39A10, 39A12.

1. Introduction. We consider ther-dimensional Euclidean spaceRr. Forx=(x1,...,

xr)T∈Rr, we define its normx =maxi∈I|xi|, whereI= {1,...,r}. InRr, we

intro-duce aconeP= {x:xi≥0, i∈I}. Then it is asolid coneinRr. It is easy to show that

P is normal, regular, minimal, strong minimal andregenerated(see Amann [3]). For two elementsxandy=(y1,...,yr)T inP, we introduce apartial ordering≤such that x< (or=)yif andonly ifxi< (or=)yifori∈Iandxymeans thatxi≤yifori∈I.

So,(Rr,)becomes apartial ordered Banach space. InRr, we also define anoperation of multiplication⊗byxy=(x1y1,...,xryr)T. In this way,(Rr,+,⊗)is apartially or-dered commutative ringby installing both this operationand the ordinary addition + with the zero element0=(0,...,0)T andthe unit elementu=(1,...,1)T. Define an ordered interval[·,·]inRr by[x,y]= {zRr:xzy}.

In ther×r-dimensional matrix spaceRr×r, we also introduce apartial ordering. If

X=(xij)r×randY =(yij)r×r are two elements inRr×r, then define thatX < (or=)Y

if andonly ifxij< (or=)yij fori,j∈Iand X≤Y means thatxij≤yij fori,j∈I.

Therefore,Rr×r also becomes apartially ordered Banach space.

Consider the following systems of nonlinear Volterra difference equations of popu-lation model with diffusion and infinite delays:

∆2um,n=A∆2

1um−1,n+1+um,n

b−Cum,n

i=0

Dium,n−i

(1.1)

for (m,n) ×Z+(0) := {1,...,M1} × ··· × {1,...,M

s} × {0,1,...}, where ∆1 and

∆2are forwardpartial difference operators,∆2

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[7, 14, 15]),A,C > (0)r×r are diagonal matrices,bRrandb>0,u·,·∈Rris a double vector sequence (only in form),D0=(0)r×r andDi∈Rr×r fori∈Z+(0).

Together with (1.1), we consider the homogeneous Neumann boundary condition

Num−1,n+1=0 for(m,n)∈∂Ω×Z+(0) (1.2)

andthe initial condition

um,j=φm,j for(m,n)∈×Z−(0):=×{...,−1,0}, (1.3)

where∆N is the normal difference,∂Ωis the boundary ofΩ(see [15]) andφm,j∈P

for(m,n)∈×Z−(0).

By asolution, we mean a double vector sequence (in form){um,n}, which is defined on(m,n)∈×Z:=×Z+(0)Z(0), satisfies (1.1), (1.2), and(1.3), respectively, when(m,n)∈×Z+(0),(m,n)∂Ω×Z+(0), and(m,j)×Z(0).

For any given initial and boundary condition (1.2) and (1.3), we can show that the ini-tial andboundary value problem (1.1), (1.2), and(1.3) have a unique solution (see [16]).

We suppose that

i=0

|Di| =D <∞,

0<φ = sup

(m,j)∈×Z−(0)φm,j<∞.

(1.4)

We write throughout this paper that

Dn= n

i=0

|Di|, δn= n

i=0

Di, D±n=Dn2±δn forn∈Z+(0). (1.5)

ThenDn,Dn±are all nonnegative, nondecreasing and bounded above byD.

SincePis regular, we can let=limn→∞D±

n. It is easy to see that

D+

n+D−n=Dn, Dn+−D−n=δn, D++D−=D, D+−D−=δ=

i=0

Di. (1.6)

Assume that

Cu> D−u. (1.7)

From Berman andPlemmons [4] or Siljak [18], we know thatC−D−is a nonsingular andinverse-positive Metzlerian matrix, i.e.,C−D−is invertible anddet(CD)−1>0. Then(C−D−)−1b>0. SinceC+δ > CD, we know, from Metzlerian matrix theory, thatC+δis invertible anddet(C+δ)−1>0.

In addition, we let

b−D+(CD)−1b>0 (1.8)

and

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It is obvious that the nonlinear Volterra difference equation of population model

∆xn=xn

b−cxn−

i=0 dixn−i

forn∈Z+(0) (1.10)

is a special case whenr=1 andwithout diffusion, where∆is the forwarddifference operator (cf. [2, 7]).

In Kocic andLadas [10], the following open problem was given.

Open problem. Obtain stability andoscillation results for (1.10).

Recently, many mathematicians have approachedsome problems of (1.10). See, e.g., Karakostas, Philos andSficas [6], Kiventidis [8], Kocic andLadas [9, 10], Kuruklis and Ladas [11], and Rodrigues [12], etc. In [14, 15, 16, 17], some problems for the nonlinear or linear Volterra difference equation of population models are considered.

It is easy to show that (1.1) has only two steady-state solutionsum,n0andum,n

(C+δ)−1b. The purpose of this paper is to give a sufficient condition for each solution of (1.1) to tendto the positive steady-state solutionum,n≡(C+δ)−1b of (1.1) by using the methodof lower andupper solutions andmonotone iterative techniques (cf. [1, 13]).

2. Some lemmas

Lemma2.1. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that{um,n}is the unique solution of (1.1), (1.2), and (1.3). Then,

um,n0,p for(m,n)∈×Z+(0). (2.1)

Proof. Consider the initial and boundary value problems

∆2vm,n≤A∆2

1vm−1,n+1+vm,n

b−Cvm,n

i=0

Divm,n−i

for(m,n)∈×Z+(0),

Nvm−1,n+1=00 for(m,n)∈∂Ω×Z+(0), (2.2)

vm,j=0≤φm,j for(m,n)∈×Z−(0)

and

∆2wm,n≥A∆2

1wm−1,n+1+wm,n

b−Cwm,n

i=0

Diwm,n−i

for(m,n)∈×Z+(0),

Nwm−1,n+1=00 for(m,n)∈∂Ω×Z+(0), (2.3)

wm,j=p≥φm,j for(m,n)∈×Z−(0).

Since

b−(C+δ)pb−Cp+D−p=b(CD)p0, (2.4)

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Lemma2.2. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that {p(n1)}is the unique solution of the Cauchy problem

p(1)

n =p(n1)⊗b−Cpn(1)+D−p forn∈Z+(0), p(j1)=p forj∈Z−(0).

(2.5)

Then{p(n1)}is nonincreasing and p(1)

n ∈C−1b+D−p ,p forn∈Z+(0). (2.6)

Proof. Consider the Cauchy problems

v(1)

n v(n1)⊗b−Cvn(1)+D−p forn∈Z+(0), v(j1)=C−1(b+D−p)≤p forj∈Z−(0),

(2.7)

and

w(1)

n w(n1)⊗

b−Cw(1)

n +D−p forn∈Z+(0),

w(j1)=pp forj∈Z−(0). (2.8)

It is easy to see thatv(n1)≡C−1(b+D−p)andw(n1)≡pare, respectively, solutions of

(2.7) and(2.8), i.e., a pair of lower andupper solutions of (2.5). So, (2.6) holds. By (2.6), we have that∆p(n1)≤0. Hence,{p(n1)}is nonincreasing. The proof is thus

complete.

Lemma2.3. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that{um,n}and{p(n1)}are, respectively, the unique solutions of (1.1), (1.2), (1.3), and (2.5). Then

um,n0,p(1)

n for(m,n)∈×Z+(0). (2.9)

Proof. Letsatisfy thatJ+J=Z+(0)andJ+J= ∅, the empty set, andbe such thatDi≥(0)r×rfori∈J+andDi< (0)r×r fori∈J−. Writeδ±=i∈J±Di. Then

we must haveδ+=D+andδ=D. Hence, using (2.6) we have,

−∞

i=0

Dip(n1)i= −

i∈J+

Dip(n1)i

i∈J−

Dip(n1)i≤ −

i∈J−

Dip(n1)i≤ −δ−p=Dp,

b−Cp(1) n

i=0

Dip(n1)ib−Cp(1)

n +D−p forn∈Z+(0).

(2.10)

Therefore,wm,np(n1)is a solution of (2.3) and(2.9) holds. Thus, the proof is complete.

For the regularity ofP, we can letp(1)=lim

n→∞p(n1). By virtue of (2.5), we can obtain p(1)=C−1(b+Dp). It follows that

limsup

n→∞ maxm∈um,np

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So, for any$=($,...,$)T>0, there exist ann1>0 andann2> n1such that

um,n<p(1)+$ fornZ+n1 (2.12) and

(0)r×r≤D−−D−n−n1−1< ($)r×r forn∈Z+(n2). (2.13)

Lemma2.4. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that {p(n2)}is the unique solution of the Cauchy problem

p(2)

n =p(n2)⊗

b−Cp(2) n +D−

p1+$ +$⊗p forn∈Z+(n2),

p(j2)=p1+$ forj∈Z−(n2).

(2.14)

Thenp(n2)is nonincreasing and um,n0,p(2)

n

for(m,n)∈×Z+(n2). (2.15) Proof. If (2.15) is not true, then there exist anm3∈Ωandann3> n2such that

um,np(n2)forn2≤n < n3andm∈Ωandum3,n3>p(n2).

Letxm,n=um,np(n2). Thenxm,n0forn2≤n < n3andm∈Ωand

xm3,n3>0. (2.16)

We can derive, from (2.14), A∆2

1xm−1,n+1−∆2xm,n+ym,nxm,n=zm,n for(m,n)∈×Z+(n2), (2.17) where

ym,n=b−Cum,n

i=0

Dium,n−i−Cpn(2) for(m,n)∈×Z+(n2),

zm,n=p(2) n

D−p

1+$ +

i=0

Dium,n−i+$⊗p

for(m,n)∈×Z+(n2).

(2.18)

It is easy to show thatym,nis bounded. We can see in the following that zm,n0. Indeed, from (2.12) and (2.13), we have

i=0

Dium,n−i= −

i=0

∆δi−1 um,n−i

= − i=0 ∆D+

i−1 um,n−i+

i=0

∆D

i−1 um,n−i

n−n1−1

i=0

∆D

i−1 um,n−i+

i=n−n1

∆D

i−1 um,n−i

≤D−

n−n1−1

p1+$ +D−−D−n−n1−1 p

≤D−p

1+$ +($)r×rp.

(2.19)

So,zm,n0for(m,n)∈×Z+(n2)from (2.18) and(2.19). It follows, from (2.17), that

∆2xm,n≤A∆2

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Consider the initial and boundary problems

∆2vm,n=A∆2

1vm−1,n+1+ym,nvm,n for(m,n)∈×Z+(n2),Nvm−1,n+1=0 for(m,n)∈∂Ω×Z+(n2),

vm,n2=0 form∈

(2.21)

and

∆2xm,n≤A∆2

1xm−1,n+1+ym,nxm,n for(m,n)∈×Z+(n2),Nxm−1,n+1≤0 for(m,n)∈∂Ω×Z+(n2),

xm,n20 form∈Ω.

(2.22)

Obviously,vm,n0is the unique solution of (2.21). Comparing (2.21) with (2.22), we see thatxm,n0for(m,n)∈×Z+(n2). But, this contradicts (2.16). Therefore, (2.15) holds.

Similarly to the proof of Lemma 2.2, we can easily show thatp(n2)is nonincreasing,

which completes the proof.

Remark2.5. As a matter of fact, we can directly use the maximum principle (see Cheng [5]) to obtain the contradiction.

We can obtain from (2.14) andthe regularity ofPthat

lim

n→∞p

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n =C−1b+D−p1+$ +$⊗p . (2.23) Therefore,

limsup

n→∞ maxm∈um,n≤C

−1b+Dp

1+$ +$⊗p . (2.24) Because$is arbitrary, we have

limsup

n→∞ maxm∈um,n≤C

−1b+Dp

1 :=p2. (2.25) Define a sequence{p%}as follows:

p%=C−1b+Dp%

−1 for%∈Z+(1),

p0=p. (2.26)

Lemma2.6. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that {p%}is defined by (2.26). Then,{p%}is nonincreasing and

(C−D−)−1b0,p% for%Z+(0). (2.27)

Proof. We rewrite (2.26) as follows:

p%=C−1Dp%

−1 for%∈Z+(1). (2.28)

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Noting thatp≥(C−D−)−1b, we have from (2.26) and(2.6), thatCp

1=b+D−p

b+D−p1. Hence,p

1≥(C−D−)−1b. By induction, we obtain (2.27). This completes the proof.

BecausePis regular, we letγ=lim%→∞p(%). From (2.26), we haveγ=C−1(b+D−γ). We can solveγ=(C−D−)−1b.

Repeating the above procedure, we can show that

limsup

n→∞ maxm∈um,n≤γ. (2.29)

From (1.8), we have thatb> D+γ. So, we can select an$0>0such thatb> D++$0)

+$0⊗p.

Let0< $ < $0. By (2.19), there exist ann4> n3andann5> n4such that

um,n< γ+$ for(m,n)∈×Z+(n4), (0)r×r≤D+−Dn+−n4−1< ($)r×r forn∈Z+(n5).

(2.30)

From (1.2), Lemma 2.1, andmaximum principle (see Cheng [5]), we know that

um,n>0for(m,n)∈×Z+(0)andcan select anη >0such that min

m∈um,n52η.

Consider the Cauchy problem

qn=qnb−Cqn−D+γ+$ $p fornZ+(n5),

qj forj∈Z−(n5). (2.31)

Repeating an argument similar to the above, we can obtain that qn <um,n for (m,n)∈×Z+(n5)and

lim

n→∞qn=C

−1bD+γ+$ $ p. (2.32)

Consequently, we have

liminfn→∞ min

m∈um,n≥C

−1bD+γ (2.33)

for$ >0being arbitrary.

Define a pair of coupledsequences{µk}and{νk}as follows:

Cµk=b−D+νk−1+D−µk−1 fork∈Z+(1), Cνk=b+D−νk−1−D+µk−1 fork∈Z+(1), ν0=(C−D−)−1b, µ0=C−1bD+ν0 .

(2.34)

Lemma2.7. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that the pair of{µk}and

{νk}is defined by (2.34). Then,

[µ0,ν0]⊇[µ1,ν1]⊇ ··· ⊇[µk,νk]⊇ ··· fork∈Z+(0) (2.35)

and

lim

k→∞µk=klim→∞νk=(C+δ)

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Proof. Because

Cµ1≥b−D+ν0=Cµ0, Cν0=(CD)ν0+Dν0=b+Dν0Cν1,

Cν0≥(C−D−)ν0=bbD+ν0=Cµ0, (2.37)

we have

[µ0,ν0]⊇[µ1,ν1]. (2.38)

We can get (2.35) by induction.

By virtue of the regularity ofP, we can letµ=limk→∞µkandν=limk→∞νk. Then

we get

Cµ=b−D+ν+Dµ, =b+DνD+µ. (2.39)

Subtracting the two equalities in (2.39), we obtainC(µ−ν)=(D++D)(µν)= D(µ−ν). So,(C−D)(µ−ν)=0.

Since(C−D)ν0=(C−D+D)(CD)−1b=bD+(CD)−1b>0from (1.8), we have, from the properties of Metzlerian matrices, that det(C−D)−1>0. Therefore, µ=ν.

It follows from (2.39) thatCµ=b−D+µ+Dµ=bδµor(C+δ)µ=b. This leads to (2.36). The proof is thus complete.

Lemma2.8. Let (1.4), (1.7), (1.8), and (1.9) hold. Suppose that the pair of{µk}and

{νk}is defined by (2.34). Then,

liminfn→∞ min

m∈um,n,limsupn→∞ maxm∈um,n

∈[µk,νk] fork∈Z+(0). (2.40)

Proof. From the above, (2.40) holds fork=0. Take an$1>0 such that$1< µ0and

b> D+(ν0+$1)D(µ0$1)+2$1p. (2.41)

For0< $ < $1, there exist ann6> n5andann7> n6such that

µ0−$ <um,n< ν0+$ for(m,n)∈×Z+(n6), (0)r×r≤D−Dn−n6−1< ($)r×r forn∈Z+(n7).

(2.42)

Now, we consider the Cauchy problems

∆¯pn=¯pnb−C¯pn+D−(ν0+$)D+(µ0$)+2$p fornZ+(n7), ¯

pj=ν0+$ forj∈Z−(n7), (2.43)

and

∆¯qn=¯qnb−C¯qn−D+(ν0+$)+D(µ0$)2$p fornZ+(n7), ¯

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Similarly to the above argument, we can obtain

¯

qn<um,n<¯pn for(m,n)∈×Z+(n7), lim

n→∞¯pn=C

−1b+D(ν0+$)D+(µ0$)+2$p ,

lim

n→∞qn¯ =C

−1bD+(ν0+$)+D(µ0$)2$p .

(2.45)

Letting$→0, we see that (2.40) holds fork=1.

Again, by repeating the above process, we have that (2.40) holds.

3. Main results and remarks. Using the seven lemmas in Section 2, together with the property thatPis normal, we get the following main result.

Theorem3.1. Let (1.4), (1.7), (1.8), and (1.9) hold. Assume that{um,n}is the unique solution of (1.1), (1.2), and (1.3). Then

lim

n→∞ m∈

um,n=(C+δ)−1b. (3.1)

For (1.10), we consider the initial condition

xj=φj≥0 forj∈Z−(0). (3.2)

Then, we have the following.

Corollory3.2. Assume thatc >∞i=0|di|,0<φ =supj∈Z−(0)φj<∞and{xn} is the unique solution of (1.10) with (3.2). Then

lim

n→∞xn= b

c+∞i=0di. (3.3)

Remark3.3. It is well known that (1.1) describes the growth ofr-species alive in Ω, that the densities of ther-populations at placemandtimenisum,n, andthat the summation represents the effects of the past history on the present growth rate in mathematical ecology. Therefore, we can only consider the caseφ>0. If this is not the case, these species do not exist. The conditionφ<∞means that the densities of these species shouldbe finite in practice. Relation (3.1) means that the growth of these species goes to an equilibrium state under ordinary conditions. Equation (1.10) is the case that we do not consider the places and diffusion.

Acknowledgement. This project is supportedby the National Natural Science

Foundation of China (# 19601016).

References

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[5] S. S. Cheng,Maximum Principles for Solutions of Second Order Partial Difference Inequali-ties, Symp. on Functional Anal. andAppl. (Taiwan), Tsing Hua univ., 1980, pp. 395– 401.

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