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Modelling animal behaviour in contests: Conventions for resource allocation

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- 7 distance sensors (d0-d6)

- 2 sensors (Rl, Rr) for detecting

the resource.

- An omnidirectional distance

sensor (Rd) and a contact

sensor (Ro) for detecting the

resource.

- A bumper sensor (bp) for

detecting collisions.

for detecting the opponent.

- Sensors for fighting ability (A)

and energy level (E).

Arena boundary

Resource

Agent 1

Agent 2

d0

d3

d6

Rl

Rr

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CBRÕRLrLhAD<r:2<>RHJL>RÓMRÑr:MQ(HwA&GRÑ>RCf<6<]ÙkÔGRL_NMØ&×-@w<r:2<>AÀNMC(ÛEü_ÐERdÑr:2<>R¿N2Ø)CBNMCBéŒLh<>Ñr@EՑ<r@BÑY:2Q“×u@wé

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(6)

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HwR<rRÑr×AÀCBRTHÕNfCBCBRÕ<>AÀNMCEL~:2CEHÔE:2ÑY:2×.R<rRÑYLÛb!àR+Ý6ADQÀQfåEÑrLh<~L>@B×.×½:2ÑrAD÷R’ÚMRCBRÑY:2Q2åECEHwAÀCBÚfL

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0.65

0.7

0.75

0.8

0.85

0.9

0.95

1

-20 -10

0

10 20 30 40 50 60

Possession rate

Ž

Mean fitness

0

0.05

0.1

0.15

0.2

0.25

0.3

-20 -10

0

10 20 30 40 50 60

Serious injury rate

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Mean fitness

SUT)V ·E·†‘J’“|³”

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(7)

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<>ÐE:2<6åEÚMÐK<>AÀCBÚ.ÐE:fL_PnRRC×.ADCBAÀ×.AD÷RH²Û

0.5

0.55

0.6

0.65

0.7

0.75

0.8

0.85

0.9

0.95

-20 -10

0

10 20 30 40 50 60

Prop. possessions by stronger

¥

Mean fitness

0.5

0.55

0.6

0.65

0.7

0.75

0.8

0.85

0.9

0.95

0.4

0.5

0.6

0.7

0.8

0.9

1

Prop. possessions by stronger

¥

Prop. peaceful possessions

SUT)V

·¦E·§‘J’“¦³”

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WrijZ]ƒfWr[og‚WYX=ZovwWri¹Xf‚dsKgbg‚W‘s†a>€KgZ‹Z]ƒf[]WrW_cw`TijX=Z]p‹i¹X-Z]i¹bdW_gqr[]`p\poZ]WYXu[]eKXKpY¼k}MWrW¨“ij‚Tef[]W”^À`[

gX.WrMcKl¹gXkgZ]ij`X.`^²pŒ{Mbvw`Tljp/eKp\W‘sB¼

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:2CEH½ADزåEÚfÐf<rADCEÚu:2PBAÀQÀA¹<]Ù.ÐE:fL‹PGRÕNf×.R@BCBAÀ×.ÔnNfÑh<Y:2CK<Öf<>ÐBRCÝ6ÐE:2<o×.ADÚfÐK<oPGRÐE:MÔBÔnRCBAÀCBÚKô

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‰ ŠX.^agqrZoZ]ƒKW6Z\[]eKW6Z]ƒKWr`[]WrZ]ijq‘gl²`TcfZ]ijbeKb ijX½`TeM[+p\ijb8efl¹gZ]ij`TXijp'¬T”2¼EusMeKW_Z]`dZ]ƒKWvw`XMefp’^À`[

(8)

÷RÑrN.AÀL_ÑrR:fLhNfCE:2PEQDÙ½ÑrRÔBÑrRL>RCK<Y:=<>AÀÓMRN2Ø&<>ÐBR¿<rRCÑr@BCELÛ

0

0.2

0.4

0.6

0.8

1

1

10

100

1000

10000 100000

Rate

­

Generations

Possession

Serious injury

Peaceful possession

0

0.2

0.4

0.6

0.8

1

1

10

100

1000

10000 100000

Rate

­

Generations

Possession by stronger

Flight

SUT)V ·®G·†‘J’“|³”

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•

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(9)

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outputs

Motor

Sensory

inputs

}

LF

LB

RB

RF

}

d3

d6

Rr

neuron

neuron

"Straight-ahead"

"Right turn"

neuron

"Flee"

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